E. annae is characterized, i.a., by its spreading, linear, obtuse leaves, which like the branches are often covered with resin, and its glabrous achenes. E. oligoglossus and E. nodosus are similar in habit but differ, i.a., by the villous achenes and different leaf-shapes. E. petraeus (see below) conies very close to E. annae but is distinguished by the distinctly ciliolate short leaves. The leaf-margins of E. annae are quite smooth or in a few specimens seen very sparsely and minutely denticulate. E. annae is said to be used in Lesotho as a medicine and for preparation of charms.
Two chromosome numbers were found in E. oligoglossus, viz., 2n = 20 and 40, respectively. I have had one collection of each in cultivation, and they are distinguishable morphologically on a number of quantitative characters. The tetraploid has larger and more distantly spaced leaves, longer and somewhat thicker peduncles, bigger flower-heads, longer involucral bracts, larger disc-florets with longer anthers, and bigger pollen grains. The same differences are observable in the material collected in nature. The herbarium material available of E. oligoglossus could be sorted on the same characters with reasonable accuracy into two form series, probably representing two cytotypes. The two putative cytotypes are treated as subspecies in the present work. Although they are likely to be effectively isolated genetically, the morphological differences are small and of a purely quantitative nature. The two subspecies have different distribution ranges and possibly also a somewhat differing ecology. Further studies are needed, however, to clarify the variation, cytology, distribution, ecology, and interrelationship of the two subspecies.
Moderately to much-branched, ± lax shrub, 0.4-1.5 m tall, sometimes sparsely woolly in axils. Leaves lanceolate to linear, spreading, 5-15 mm long. Flowerheads solitary in upper axils forming racemes or corymbs, on short peduncles 5-30 mm long, radiate, yellow; involucre hemispherical, 3-7 mm diam., bracts 5 or 6, connate < 1/4. Cypselas densely white-hairy, mucilaginous when wet.
Shrub, 0.2-1.5 m high; dense, rounded, erect, resiniferous, glabrous; stems closely leafy, later nude with leaf scars. Leaves alternate, sessile, closely set, ± spreading; blade linear, 5-20 x 1-2 mm, apex obtuse, minutely apiculate, base dilated, ± half-clasping, margins entire, smooth; thick, leathery. Heads radiate, solitary; peduncles 5-25 mm long, lateral in apical leaf axils, in a corymb or ± raceme. Involucral bracts uniseriate, 5 or 6, connate to 1/4-1/3, veins absent to many-veined, ecalyculate. Flowers: ray florets 2-5, female, fertile; disc florets 8-20, bisexual; all florets yellow; Feb.-Apr. Fruit with cypsela oblong-obovate, glabrous, brown, ribs 5-8. Pappus of many barbellate bristles, white.
Lax shrub to 1.5 m, sometimes sparsely woolly in axils. Leaves lanceolate to linear, spreading, 5-15 mm long. Flower heads radiate, solitary on short peduncles in upper axils, yellow. Achenes densely white-hairy, mucilaginous when wet.
An erect much-branched shrub, 0.2-1.5 m high, rather dense and rounded, strongly resiniferous and normally with rich surface resin, wholly glabrous or seldom sparsely arachnose in the upper leaf-axils. Young branches closely leafy, becoming nude and marked with scalelike leaf-scars. Leaves closely set, ± spreading (erecto-patent when young), linear, entire, 0.5-2 cm long, 1-2 mm wide, thick and coriaceous, but not markedly rugulose when dry, semiterete or flattened, obtuse and minutely apiculate with a whitish point, with a minutely muricate surface, bright green or glaucous, often with surface resin; adaxial side flat or slightly canaliculate with a narrow impressed midvein; abaxial side convex or somewhat keeled, midribbed; leaf-base dilated, almost half-clasping. Peduncles lateral in the leaf-axils, in a corymb- or shortly raceme-like arrangement at the branch ends, 0.5-2.5 cm long, slender, 0.2-0.5 mm thick, terete, faintly striate. Peduncle-base surrounded by two lateral reduced leaves; these subulate, 0.5-1.5 mm long. Involucre campanulate with a narrow, often somewhat cuneiform base, 3-7 mm wide and 3-6 mm high. Involucral bracts uniseriate, 5(-6), connate to 1/4-1/3 or sometimes almost to the middle, smooth or midveined, sometimes 3-many-veined, herbaceous or somewhat coriaceous with membranous margins, lanceolate-ovate, 3-7 mm long, 1-2.5 mm wide, acute or acuminate. Receptacle flat or somewhat convex, alveolate. Ray-florets 2-5, yellow. Tube 1-1.5 mm long, cylindrical. Lamina 2.5-5 mm long, 1-2.5 mm wide, oblong-elliptic-oblong, 4(-9)-veined, acute-truncate. Style terete; style branches 0.5-1 mm long. Disc-florets 8-20, often 13. Corolla 3.5-4 mm long. Tube 1-1.5 mm long, cylindrical. Limb narrowly campanulate, 2-3 mm long; lobes deltoid, 0.5-0.7 mm long. Anthers 1.3-1.8 mm long incl. the ovate-narrowly ovate subacute appendage. Style terete with swollen base; stylophore short; style branches 0.3-0.7 mm long with truncate-slightly convex tips. Pappus bristles numerous, 1-2 mm long, erect, white. Achenes oblong-obovate, 2-2.5 mm long, 0.8-1.5 mm wide, glabrous, light or yellowish brown, with 5(-8) prominent resiniferous ribs.
Dry Grassland; Dwarf Karroid Shrubland. Grassland, karoo, dominating disturbed veld.
It is a montane species, growing at altitudes between 1500 and 2700 m s.m. The habitat is various types of grassveld (e.g. Karroid Danthonia Mountain Veld, Themeda-Festuca Alpine Veld) or False Karroo. In disturbed veld the species may play a dominating role.
Roggeveld to Hex River Mountains and Matjiesfontein to Lesotho.