Differing from subspecies uncinactumn by the predominantly 5-lobed leaf with lobes narrow, more cuneate at the base, often trifid or deeply serrate with acumi- nate teeth (Fig. 1, e-k) ; pedicels and usually also rachises of infloreseence pubescent with straight and spreading hairs (Fig. 3, c); helmet mostly rounded above the very slight beak, or often conic and nearly vertical above the distinct beak (Fig. 4, d, top).
Shaded ravines, damp slopes, stream banks and springheads of rich deciduous woods, or occasionally in more open, weedy areas. Mountains from southwestern Pennsylvania and western Maryland to those of eastern Tennessee and north- western South Carolina; also in disjunct areas in the inner Coastal Plain and eastern Piedmont of southeastern Virginia and North Carolina (Fig. 6).