Trees to 30 m tall. Petiole 1-2.5 cm; leaf blade oblong-ovate, oblong-lanceolate, or lanceolate, 10-23 cm, abaxially covered with yellowish brown scalelike glands and sparsely pilose along veins when young, glabrescent, base rounded to broadly cuneate but narrowly cuneate when young, margin with bristlelike teeth 2-4 mm, apex long acuminate. Male inflorescences 5-16 cm. Female flowers 1(-3) per cupule. Cupules on a short spike, 2.5-3.5 cm in diam. including bracts, covered with slightly pubescent spinelike bracts. Nut 1 per cupule, globose-ovoid, 1.5-2 cm, longer than wide. Fl. May-Jul, fr. Sep-Oct.