Croton macrostachyus Hochst. ex Delile
  • Voy. Abyssinie


Cite taxon page as 'WFO (2020): Croton macrostachyus Hochst. ex Delile. Published on the Internet;http://www.worldfloraonline.org/taxon/wfo-0000931591. Accessed on: 10 Jul 2020'

Morphology

A tree, in savannah, up to 30–50 ft high

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    Flowers white

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    Fruit slightly 3-lobed, about 1/2 in. diam.

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    Bark light or dark grey, finely to deeply longitudinally fissured.

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    A dioecious or sometimes monoecious shrub or spreading tree up to 25 m. tall, although more commonly 6–12 m. tall.

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    Male flowers: pedicels 4–10(–14) mm. long, densely stellate-pubescent; sepals 5, ovate, 3 mm. long, 2.5 mm. broad, obtuse, stellate-pubescent without, pubescent within at the middle and towards the apex, pale green; petals 5, oblanceolate-oblong, 3.5 mm. long, 1.5–2 mm. wide, obtuse, ± glabrous without, densely villous within and on the margin, pale yellow; disc-glands rounded, hairy; stamens 15–20, filaments 4 mm. long, villous in the lower half, anthers 1 mm. long; receptacle densely villous.

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    Racemes terminal, (7–)15–32 cm. long, up to 3 cm. wide, either all ♂ or all ♀, or mostly ♂ with a few ♀ flowers at the base, or sometimes half ♂ and half ♀ or mostly ♀ with a few ♂ flowers at the top, the ♂ flowers in clusters along the axis, the ♀’s usually solitary within the bracts, or with 1 ♀ flower accompanied by 1 or more ♂ flowers per cluster; axis densely stellate-strigose, or evenly or sparingly stellate-pubescent; bracts linear, 2–3(–5) mm. long, sparingly pubescent or subglabrous.

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    Fruit subtrilobed, rarely 4-lobed, often more particularly so at the apex, 8–9 mm. long, 8–10(–15) mm. wide, loculicidal, evenly stellate-pubescent-sublepidote.

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    Female flowers: pedicels 2–5(–8) mm. long, stouter than the ♂ pedicels, scarcely elongating in fruit, densely fulvous stellate-pubescent; sepals 5, lanceolate or ovate-lanceolate, 3 mm. long, 1.5 mm. broad, obtuse, evenly stellate-pubescent without, sparingly stellate-pilose within; petals 0, or, if present, linear and smaller than in the ♂ flowers; staminodes 0–1(–2), filament-like; disc-glands free, ± truncate; ovary trigonous, 2 mm. diameter, densely fulvous adpressed stellate-pubescent-sublepidote; styles 3(–4), spreading, 2.5 mm. long, pubescent at the base, otherwise glabrous, deeply bipartite, the lobes linear, sometimes bifid at the apex, 2 mm. long.

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    Older twigs greyish brown.

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    Young twigs sparsely, evenly or densely greyish ± adpressed stellate-pubescent, the hairs often with a projecting central ray.

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    Stipules linear or linear-subulate, 5–14 mm. long, entire or with a few subulate lateral lobes, sparingly stellate-pubescent or subglabrous, caducous.

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    Petioles 1.5–11 cm. long, usually sparingly stellate-pubescent, sometimes either densely so or almost glabrous; leaf-blade broadly ovate to elliptic-ovate, 4–15(–20) cm. long, 2.5–10(–13) cm. wide, obtusely or subacutely acuminate, rounded or cordate, with 2–4(–8) subsessile or stipitate discoid basal glands visible or not from above, crenulate-serrulate to subentire, thinly to firmly chartaceous, 5–7(–9)-nerved from the base, remaining lateral nerves 4–8(–11) pairs, usually fairly prominent beneath, densely fulvous (above) and greyish (beneath) stellate-pubescent at first, later becoming evenly or sparingly adpressed stellate-pubescent and dark green above, and evenly or sparingly greyish hirsute-stellate-pubescent beneath.

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    Seeds ellipsoid, 7 mm. long, 4 mm. wide, longitudinally rugulose, grey, somewhat shiny, with a large waxy caruncle 4.5 mm. long and 4 mm. wide.

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    Female flowers: pedicels 2–5 mm long, not or scarcely extending in fruit, stout, densely stellate-pubescent; sepals 5, 3–3.5 × 1.5–2 mm, ovate-lanceolate, evenly stellate-lepidote without, sparingly stellate-pubescent within, grey-green; petals 0 or 5, 0.5–1 mm long, linear to subulate, subglabrous; disk 5-lobed, the lobes truncate, pubescent; ovary 2 mm in diameter, trigonous or quadrangular, densely lepidote; styles 3–4, 3 mm long, spreading, 2-partite with the segments filiform-linear, minutely puberulous.

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    Fruits 0.8–1 × 0.8–1.5 cm, trilobate or quadrilobate, loculicidal or septicidal, densely to evenly scurfily stellate-pubescent, pale greyish-green.

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    Seeds 7 × 4 mm, ellipsoid, rugulose, grey; caruncle 4.5 × 4 mm, waxy.

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    Racemes up to 35 cm long, terminal, usually androgynous, sometimes male or female; bracts shorter than the stipules but otherwise resembling the stipules.

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    Flowers fragrant.

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    Male flowers: pedicels 0.5–1 cm long; sepals 5, 3 × 2–2.5 mm, ovate, stellate-lepidote without, pubescent within, pale green; petals 5, 3–3.5 × 1.5–2 mm, oblanceolate-oblong, subglabrous without, villous within and on the margin, pale creamy-yellow in colour; disk glands rounded, pilose; stamens 15–20, filaments 4 mm long, villous below, anthers c. 1 mm long; receptacle densely villous.

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    Petioles 3–12 cm long.

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    Leaf blades 6–18 × 4–14 cm, ovate, acuminate at the apex, shallowly crenate-serrate to subentire on the margins, cordate or subcordate at the base with a pair of shortly stipitate to subsessile basal discoid glands on the lower surface, chartaceous, sparingly appressed stellate-pubescent to fimbriate-lepidote above, and sparingly to densely so or patent stellate-tomentose beneath; 5–7-nerved from the base, lateral nerves in 4–9 pairs, ± prominent beneath.

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    Young twigs, petioles and inflorescence axes densely, evenly or sparingly greyish appressed or patent stellate-pubescent, later glabrescent and becoming dark grey-brown and lenticellate.

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    Stipules 5–7 mm long, linear to filiform-setaceous, soon falling.

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    Bole up to 1.8 m in circumference.

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    Bark smooth to closely reticulate, pale grey or greyish-brown, slightly corky.

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    A tree up to 30 m tall, dioecious or sometimes monoecious; crown much-branched, spreading, rounded.

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      Bibliography

     Information From

    Euphorbiaceae
    • A CC0 1.0 Universal (CC0 1.0).
    Plants Of the World Online Portal - FWTA
    https://www.kew.org/science/who-we-are-and-what-we-do/strategic-outputs-2020/plants-of-the-world-online
    http://www.plantsoftheworldonline.org/terms-and-conditions
    • B The Trustees of the Royal Botanic Gardens, Kew
    Plants Of the World Online Portal - FZ
    https://www.kew.org/science/who-we-are-and-what-we-do/strategic-outputs-2020/plants-of-the-world-online
    http://www.plantsoftheworldonline.org/terms-and-conditions
    • C
    Plants Of the World Online Portal - FTEA
    https://www.kew.org/science/who-we-are-and-what-we-do/strategic-outputs-2020/plants-of-the-world-online
    http://www.plantsoftheworldonline.org/terms-and-conditions
    • D
    World Flora Online consortium
    http://www.worldfloraonline.org/organisation/WFO
    World Flora Online Data. 2018.
    • E CC0 1.0 Universal (CC0 1.0).
    Global Tree Assessment
    http://www.bgci.org/
    BGCI. 2018. GlobalTreeSearch online database. Botanic Gardens Conservation International. Richmond, U.K. Available at www.bgci.org. Accessed on 30/11/2018.
    • F Botanic Gardens Conservation International