Tree ca 5-30 m high, with trunk 0.7-6 dm thick; monoecious; young twigs + scurfy with compact dendritic hairs. Leaves membranous or chartaceous; petioles scurfy 3-20 cm long, the glands at apex of petiole 2-6 (plus a cluster of smaller ones usually present on ventral side), patelliform, more or less stalked, 0.5-1.2 mm across (stalk usually about equaling diam of gland); stipules linear-lanceolate to subulate, 3-10 mm long; blades mostly ovate, (7-) 10-30 cm long, (2.5-) 7-23 cm broad, sparsely pubescent above with scattered dendritic and hispid hairs (eventually glabrate), sparsely to densely tomentose beneath with stellate and dendritic hairs (these multiradiate, mostly 0.3-0.6 mm wide) and minutely glandular-dotted, 5-7-nerved at base, with ca 6-10 additional veins on either side, the veinlet reticulum promi- nent beneath, the base truncate to deeply cordate, the margins subentire to ob- scurely or rather prominently denticulate, the apex rather abruptly acuminate or caudate. Inflorescences terminal, usually bisexual, racemiform, (7-) 20-40 cm long, typically with the 5-20 proximal cymules I, the distal (and sometimes the first proximal) cymules j; bracts triangular or lanceolate, tomentose, inconspicuous, ca 1-1.5 mm long, subtending several flowers. Staminate flowers with stellate- tomentose pedicels ca 5-8 mm long; calyx-lobes 5, stellate-tomentose, triangular, 2-2.2 mm long; receptacle densely villose; petals obovate, 1.9-2.5 mm long, villose especially on the margins; stamens (13-) 14-18(-20), the filaments glabrous (except at base), 2-2.8 mm long, the anthers elliptic, 0.6-0.8 mm long. Pistillate flowers with stout stellate-tomentose pedicels (2-) 3-6 mm long; calyx-segments 5, whitish- or yellowish-tomentose outside, dark brownish inside, elliptic to oblong, more or less valvale, 2.3-2.8 mm long, 1.3-1.8 mm broad; disc 2-3 mm wide, notched or lobed into 5 entire or toothed segments; petals filiform-subulate to rarely narrowly ligulate, 0.5-2.1 mm long, 0.1(-0.4) mm broad; ovary orange- or brownish-tomen- tose with stellate and dendritic hairs (hispid radii ca 0.3-0.6 mm long), the styles free, bifid or bipartite, glabrous or rarely sparsely stellate, at the base, (2.5-)3-5 mm long, the slender branches mostly 2.5-3 mm long. Capsules subglobose, yellowish- brown, stellate-hispidulous, ca 6.5 mm diam, the columella slender, 3.3-3.5 mm long; seeds plump, brown, obscurely costate-roughened, 3.1-3.9 mm long, 2.6-2.8 mm broad, the caruncle ovate, ca 1 mm wide.