Trees to 20 m tall; bark gray-brown, smooth. Branchlets dark gray, angular, densely gray pubescent when young, glabrescent. Buds stipitate, with 2 scales, sparsely pubescent. Petiole 1.5-5.5 cm, densely pubescent; leaf blade suborbicular, rarely broadly ovate, 4-9 × 2.5-9 cm, abaxially light green or glaucescent, densely or sparsely brown hispidulous, rarely subglabrous, sometimes bearded in axils of lateral veins, adaxially dark green, sparsely villous, base rounded or broadly cuneate, rarely cuneate or subcordate, margin undulate-serrate, apex rounded, rarely acute; lateral veins 5-10 on each side of midvein. Female inflorescences 2-8 in a raceme, subglobose or oblong, 1-2 cm; peduncle 2-3 mm; bracts 3-4 mm, woody, base cuneate, apex rounded, 5-lobed. Nutlet broadly ovate, ca. 3 mm, with papery wings ca. 1/4 as wide as nutlet. Fl. May-Jul, fr. Jul-Aug.