Lycopodium clavatum L.
  • Sp. Pl.
  • Stag’s-horn clubmoss


Cite taxon page as 'WFO (2021): Lycopodium clavatum L. Published on the Internet;http://www.worldfloraonline.org/taxon/wfo-0001107341. Accessed on: 26 Sep 2021'

General Information

1. Lycopodium clavatum L., Sp. Pl. 1101 (1753). Lectotipo (designado por Jonsell y Jarvis en Jarvis et al., 1993): Herb. Burser XX: 49 (UPS).

Por B. Øllgaard.

Rizoma principal generalmente epigeo; últimas ramitas teretes, isofilas; hojas 6-8(-10) x 0.5-0.8 mm, lineares a aciculares, enteras a esparcidamente denticuladas, terminadas en un pelo incoloro o en un ápice membranáceo; estróbilos sésiles o pedunculados; esporofilos subpeltados, el pedículo con un ala media, basiscópica, membranácea conectando el pliegue basiscópico de la cara exterior al eje del estróbilo; células epidérmicas del esporangio con paredes laterales fuertemente sinuadas, con invaginaciones y evaginaciones en forma de bolsas; esporas reticuladas en todas sus caras. Claros y hábitats expuestos en bosques montanos altos, páramos. Ch, G-P. 1000-3800 m. (Cosmopolita, con excepción de Australia.)

Japón tiene tetraploides con estróbilos más agrupados que los diploides. Hay formas similares neotropicales pero se desconoce su citología.

  • Provided by: [G].Flora Mesoamericana
    • Source: [
    • 1
    • 7
    • ]. 

    Stolons subterranean, slender and creeping, 1 or 2 times forked, green, with sparse entire leaves; lateral branches erect, 20-25 cm tall, 3-5 times forked, sparse, flattened (young branches terete), stem together with leaves 9-12 mm in diam. Leaves spirally arranged, dense, angled upward, lanceolate, 4-6 × ca. 1 mm, leathery, with transparent hairs, midrib visible on both surfaces, base cuneate, decurrent, sessile, margin entire, apex acuminate. Strobili 2(or 3) together on a peduncle, peduncle up to 12 cm, bracts of peduncle spirally arranged, sparse, herbaceous, narrowly lanceolate; strobili erect, terete, 3.5-4.5 cm × ca. 4 mm, subsessile or with short, equally long pedicels; sporophyll broadly ovate, ca. 1.5 × 1.3 mm, papery, margin membranous, erose, apex acute, with short acute tip. Sporangia slightly exposed.

  • Provided by: [H].Flora Of CHina @ efloras.org
    • Source: [
    • 5
    • ]. 

    Horizontal stems on substrate surface. Upright shoots clustered, 0.6--1.2 cm diam., dominant main shoot with 3--6 branches mostly in lower 1/2. Lateral branchlets few and like upright shoots; annual bud constrictions abrupt, branchlets mostly spreading. Leaves spreading, often somewhat ascending in distal 1/3 of branches, medium green, linear, 4--6 X 0.4--0.8 mm; margins entire; apex with narrow hair tip 2.5--4 mm. Peduncles 3.5--12.5 cm, with remote pseudowhorls of appressed leaves, loosely branched into 2--5 alternate stalks, 0.5--0.8 cm. Strobili 2--5 on alternate stalks (if double, usually with stalks 5--8 mm), 15--25 X 3--6 mm. Sporophylls 1.5--2.5 mm, apex abruptly reduced to hair tip. 2 n = 68.

  • Provided by: [F].Flora of North America @ efloras.org
    • Source: [
    • 6
    • ]. 

    Lycopodium clavatum L. ssp. clavatum, Sp. Pl. 1101. 1753. Fig. 81 A–D.Tallos 1–3 mm de diámetro; ramifi­caciones late­rales erectas, rígidas o más o menos de­cumben­tes, 10–40 cm de largo, sub­dicotómicas o pinnadamente divididas; hojas adpresas a patentes, 4–7 mm de largo y 0.5–1 mm de ancho, ate­nua­das dis­talmente, denti­culadas o con una seta apical de 1–3 mm de largo, la seta blanquecina; estróbilos cilín­dri­cos, 2–6.5 cm de largo, 1–3 por pedículo, erectos o leve­mente curvos; esporofilos delta­do-ovados, 2–3.5 mm de largo y 0.8–2 mm de ancho, márgenes erosos.Áreas alteradas; Croat 43083, Rueda 12527;800–1550 m; Canadá a Para­guay y Brasil, las Antillas, Europa, Asia e Islas del Pacífico. VU

  • Provided by: [E].Flora de Nicaragua
    • Source: [
    • 8
    • ]. 

    "Horizontal stems superficial, elongate, arching and rooting at intervals, 2–3 mm thick, not dorsiventral, the lvs curved upwards; erect stems branched, to 2.5 dm; lvs mostly 10-ranked, uniform, linear-subulate, 4–7 × 0.5–0.8 mm, slightly or scarcely toothed, attenuate to a hyaline hair-like tip 2–3 mm; cones 1.5–11 cm, 3–7 mm thick, 1–6 on remotely bracteate peduncles 1.5–15 cm; sporophylls yellowish, broad-based, abruptly acuminate, with scarious erose margins; sporangia reniform, ca 1.5 mm wide; spores 28–36 microns in diameter, the sides convex, the commissural and outer faces angular-reticulate, the commissures not in furrows; 2n=68. Open dry woods and rocky places in acid soil; circumboreal, s. in e. U.S. to N.C., W.Va., Mich., and Io."

  • Provided by: [M].New York Botanical Garden
    • Source: [
    • 9
    • ]. 

    Morphology

    Tiges principales épigées, pouvant atteindre 4 m de long et 3 mm d'épaisseur, rampant sur le sol ou parfois s'appuyant sur des sousarbustes, émettant des rameaux ascendants à dressés atteignant 15-30 cm de long.' Feuilles de 4-8 mm de long sur 0,5-0,75 mm de large, disposées en spirale ou rarement subverticillées, celles des tiges couchées toutes semblables, linéaires-subulées, à bords entiers à irrégulièrement denticulés, aiguës et à sommet prolongé par une soie longue de 2-4 mm. Epis de 1,5-7 cm de long sur 3-5 mm de diam., par 2-5 au sommet d'un pédoncule long de 5-19 cm, dressé, portant des feuilles pouvant atteindre 5 mm de long, espacées, alternes à subverticillées; pédicelles de 0-25 mm de long; bractées de 2-5 mm de long sur 1,75-2 mm de large, subpeltées, ovales à triangulaires, à bords érodés-denticulés, longuement acuminées en une soie longue de 2 mm.

  • Provided by: [D].Flore d'Afrique Centrale
    • Source: [
    • 10
    • ]. 

    Triebe bis über 1 m lang, kriechend, mit 5–15 cm hohen, aufstrebenden Ästen. Blätter 3–6 mm lang und 0,5–1 mm breit, vielreihig, spiralig angeordnet, in eine weisse Haarspitze auslaufend. Sporangienähren zu 2–3 auf langem, locker beblättertem Stiel

  • Provided by: [B].Flora Helvetica – Illustrierte Flora der Schweiz
    • Source: [
    • 11
    • ]. 

    Sporophylls different from foliage leaves, grouped into pedunculate strobili. Horizontal stem producing much-branched, erect stems at intervals. Leaves hair-pointed. Erect stems not laterally compressed.

  • Provided by: [A].e-Flora of South Africa
    • Source: [
    • 12
    • ]. 

    Terrestrial. Stems erect, branching from a prostrate, branched main stem, 80-300 mm long. Leaves erect from branches at right angles, with aristate tips bent forward, 3-5 mm long, up to 0.6 mm wide; sporophylls ephemeral, distinct from sterile leaves. Sporangia short-stalked, borne in sporophyll axils in a terminal strobilus.

  • Provided by: [A].e-Flora of South Africa
    • Source: [
    • 13
    • ]. 

    Plants terrestrial. Stems wide-creeping, rooting at irregular intervals, anisotomously branched, main stems with indeterminate growth, to 2 mm in diameter, isophyllous, lateral branches erect, arising in a dorsi lateral position on the main stem, isophyllous, fertile branches erect, anisotomously branched, sparsely leafy, isophyllous, heterophyllous, to 180 mm long. Leaves herbaceous, acicular, entire, aristate, patent to imbricate, to 7 mm long, to 1 mm wide. Strobili terminal, in groups of 3-5, to 30 mm long, isophyllous, sporophylls ephemeral, herbaceous, subpeltate, often with a basiscopic membranous wing on the stalk, narrowly to broadly ovate, aristate, laciniate, to 4.5 mm long, to 2 mm wide. Sporangium reniform, short-stalked, borne adaxially near the sporophyll base, isovalvate, to 1.3 mm in diameter. Spores spheroidal to subtriangular, trilete, reticulate, (34-)37.5(-38) µm in equatorial diameter, (30-)31 (-34) µm in polar diameter.

  • Provided by: [A].e-Flora of South Africa
    • Source: [
    • 14
    • ]. 

    Perennial with prostrate main stem and dichotomously branched, erect stems. Leaves with a translucent hair point. Strobili pedunculate, in groups of 2-5.

  • Provided by: [A].e-Flora of South Africa
    • Source: [
    • 15
    • ]. 

    Terrestrial. Main stem creeping, up to 3 mm in diameter, producing dichotomously branched upright stems at 50-100 mm intervals. Leaves narrowly lanceolate, patent to suberect or imbricate, up to 5 x 0.5 mm, with a translucent hair point up to 3 mm long apically, margin usually entire but sometimes ciliate in leaves on main stem. Strobili pedunculate, 20-80 mm long in groups of 2-5 (rarely solitary) at apex of sparsely leafy peduncle up to 190 mm long. Sporophylls broadly ovate, acuminate, 2-3 x 1.5-2 mm, margin finely lacerate.

  • Provided by: [A].e-Flora of South Africa
    • Source: [
    • 16
    • ]. 

    Sürünücü gövde genellikle toprak altında, kökler toprağa ilk temas eden noktalardan çıkar, 1-2 kez çatallanır, 2 metreye kadar uzayabilir. Topraküstü sürgünler genellikle dik, 9-25 cm boyunda, 3-5 kez çatallanmış, düzleşmiş (bazen genç sürgünlerde sütunsu), gövde yapraklarla birlikte 7-12 mm çapında. Yapraklar helezonî dizili, sık, üste doğru gövdeden dar açı ile uzaklaşır, mızraksı 4-6 × 0,8-1 mm, sert, üzerinde saydam tüyler bulunur; ana damar her iki yüzde de belirgin, tabana doğru daralıcı, kenarları gövdeye doğru dekürent, sapsız, kenarlar dümdüz ya da belli belirsiz birkaç dişli, uç kısım sivri, yaprak kadar uzun olabilen tüysü çıkıntılı. Kozalak dik, sütunsu, 13-45 × 3-4 mm, 4-12 cm uzunluğundaki bir sap üzerinde 2(- 3) tane, sapın bırakteleri helezonî, otsu, darca mızraksı. Spor yaprakları genişçe yumurtamsı 1,5 × 1,3 mm, ince yapılı, kenarları zarımsı, düzensiz dişli, ucu sivri, kuyruklu. Spor keseleri az miktarda genişlemiştir. Spor çok sayıda.

  • Provided by: [N].Resimli Türkiye Florası (the Illustrated Flora of Turkey)
    • Ecology

      7–9. Moosige Wälder und Weiden, (kollin-)montan-subalpin. A, M, selten J.

    • Provided by: [B].Flora Helvetica – Illustrierte Flora der Schweiz
      • Source: [
      • 11
      • ]. 

      Habitat

      Mesic Grassland. Moist or seasonally moist habitats, especially road cuttings and sheltered mountain slopes amongst rocks.

    • Provided by: [A].e-Flora of South Africa
      • Source: [
      • 13
      • ]. 

      Savanes, forêts claires, lisières de galeries forestières, talus de routes, en sites souvent humides et sablonneux; de (800-)1500 à 3350 m d'altitude dans l'aire de la Flore.

    • Provided by: [D].Flore d'Afrique Centrale
      • Source: [
      • 10
      • ]. 

      Forest margins, sheltered mountain slopes amongst rocks.

    • Provided by: [A].e-Flora of South Africa
      • Source: [
      • 12
      • ]. 

      Terrestrial, on moist or seasonally moist slopes in grasslands, forest margins, and disturbed or man-made habitats such as road cuttings, growing in exposed or partially shaded conditions. Not edaphically bound.

    • Provided by: [A].e-Flora of South Africa
      • Source: [
      • 14
      • ]. 

      Moist forest margins and road cuttings, 300-1500 m.

    • Provided by: [A].e-Flora of South Africa
      • Source: [
      • 15
      • ]. 

      Sheltered mountain slopes amongst rocks and continually moist margins of montane forest. 1600-2300 m.

    • Provided by: [A].e-Flora of South Africa
      • Source: [
      • 16
      • ]. 

      Distribution

      Zones tempérées et froides des deux hémisphères; montagnes de l'Asie, de l'Afrique et de la Polynésie intertropicales.

    • Provided by: [D].Flore d'Afrique Centrale
      • Source: [
      • 10
      • ]. 

      The species is widespread in sub-Saharan Africa and the western Indian Ocean region.

    • Provided by: [A].e-Flora of South Africa
      • Source: [
      • 14
      • ]. 

      Grootwinterhoek Mtns to Knysna, E and W Africa, Mascarenes.

    • Provided by: [A].e-Flora of South Africa
      • Source: [
      • 15
      • ]. 

      South-western Cape Province to Transkei, Natal, Lesotho, Swaziland, Transvaal, Zimbabwe, Mozambique, Malawi, Angola, Zaire, Tanzania, Kenya, Uganda, Ethiopia. Sudan, Cameroun, as well as Madagascar. Reunion, Mauritius and São Tomé.

    • Provided by: [A].e-Flora of South Africa
      • Source: [
      • 16
      • ]. 

      Distribution Map

       
      • Native distribution
      Found in
      • Europe Northern Europe Ireland
      • Southern America Brazil Roraima
      • Bahia
      • Paraní
      • Rio Grande do Sul
      • Santa Catarina
      • Espirito Santo
      • Minas Gerais
      • Rio de Janeiro
      • São Paulo
      • Goiás

      Synonyms

      Nationally Preferred Names

      NameLanguageCountry
      Stag’s-horn clubmossEnglishIE
      GöbektozuTurkishTR
      Garbhógach na mbeannIrishIE

      Other Local Names

      NameLanguageCountry
      Lycopode en massueFrenchCH
      Garbhóg (-óige) (or Garbhógach) sléibheIrishIE
      Wolf’s claw (wolfes-claw)EnglishIE
      Keulen-BärlappGermanCH
      Wandering fogEnglishIE
      Lion’s pawEnglishIE
      Common club-moss, lycopode à massue
      Patta-luf mazzacunRaeto-RomanceCH
      Licopodio clavatoItalianCH

        Bibliography

       Information From

      e-Flora of South Africa
      e-Flora of South Africa. v1.21. 2018. South African National Biodiversity Institute. http://ipt.sanbi.org.za/iptsanbi/resource?r=flora_descriptions&v=1.21
      • A All Rights Reserved
      Flora Helvetica – Illustrierte Flora der Schweiz
      http://www.flora-helvetica.ch
      Flora Helvetica – Illustrierte Flora der Schweiz, Konrad Lauber, Gerhart Wagner, Andreas Gygax. Haupt Verlag 2018.
      • B All Rights Reserved
      Brazilian Flora 2020 project - Projeto Flora do Brasil 2020
      http://floradobrasil.jbrj.gov.br
      Brazil Flora G (2014): Brazilian Flora 2020 project - Projeto Flora do Brasil 2020. v393.147. Instituto de Pesquisas Jardim Botanico do Rio de Janeiro. Dataset/Checklist. doi:10.15468/1mtkaw
      • C Group Brazil Flora, REFLORA Program
      Flore d'Afrique Centrale
      https://www.floredafriquecentrale.be
      • D http://creativecommons.org/licenses/by/4.0/ Attribution 4.0 International (CC BY 4.0)
      Flora de Nicaragua
      http://www.tropicos.org/projectwebportal.aspx?projectid=7&pagename=Home&langid=66
      WD Stevens, CU Ulloa, A Pool and OM Montiel. Missouri Botanical Garden Press, 2001
      • E Missouri Botanical Garden
      Flora of North America @ efloras.org
      http://www.efloras.org/flora_page.aspx?flora_id=1
      'Flora of North America @ eFloras (2008). Published on the Internet http://www.efloras.org/flora_page.aspx?flora_id=1 [accessed August 2016]' Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
      • F Flora of North America Association
      Flora Mesoamericana
      http://www.mobot.org/MOBOT/fm/
      Gerrit Davidse, Mario Sousa Sánchez, A. O. Chater, Universidad Nacional Autónoma de México. Instituto de Biología, Missouri Botanical Garden, Natural History Museum (London, England) UNAM, 1994
      • G Missouri Botanical Garden
      Flora Of CHina @ efloras.org
      http://www.efloras.org/flora_page.aspx?flora_id=2
      'Flora of China @ eFloras (2008). Published on the Internet http://www.efloras.org/flora_page.aspx?flora_id=2 [accessed August 2016]' Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
      • H Missouri Botanical Garden
      Irish Vernacular Names
      https://www.missouribotanicalgarden.org/plant-science/plant-science/research-staff/article/380/wyse-jackson-peter-s.aspx
      • I All Rights Reserved
      Lycopodiaceae
      World Flora Online Data. 2017.
      • J CC0 1.0 Universal (CC0 1.0).
      MBG Floras Images
      http://www.tropicos.org/ImageSearch.aspx
      Flora images. Missouri Botanical Garden. Accessed on Jun. 2018.
      • K Missouri Botanical Garden
      • L Missouri Botanical Garden
      New York Botanical Garden
      Descriptions of plants should be attributed to the full citation for each individual article, chapter or book that is the source for each record, which should include the authors of original publication.
      • M Content licensed under Creative Commons Attribution-NonCommercial-ShareAlike 3.0 Unported License
      Resimli Türkiye Florası (the Illustrated Flora of Turkey)
      http://www.turkiyeflorasi.org.tr
      Güner, A., Kandemir, A., Menemen, Y., Yıldırım, H., Aslan, S., Ekşi, G., Güner, I. ve Çimen, A.Ö. (edlr.) (2018). Resimli Türkiye Florası, Cilt 2. ANG Vakfı Nezahat Gökyiğit Botanik Bahçesi Yayınları. İstanbul.
      • N All Rights Reserved
      • O Resimli Türkiye Florası (the Illustrated Flora of Turkey).
      World Flora Online consortium
      http://www.worldfloraonline.org/organisation/WFO
      World Flora Online Data. 2018.
      • P CC0 1.0 Universal (CC0 1.0).