The World Flora Online

2. Anogramma leptophylla (L.) Link, Fil. Spec. 137 (1841).

Por R.C. Moran.

Polypodium leptophyllum L., Sp. Pl. 1092 (1753). Lectotipo (designado por Morton, 1970): País desonocido, Anon. (microficha MO! ex LINN-1251.56).

Anogramma guatemalensis (Domin) C. Chr., Gymnogramma leptophylla (L.) Desv., G. leptophylla var. mexicana M. Martens et Galeotti, Pityrogramma guatemalensis Domin.

Rizoma peloso, los tricomas articulados; hojas 5-20(-30) x 2-5 cm; pecíolo 4-15 cm, pajizo o pardo-rojizo; lámina ovada a deltada, 2-4-pinnada; segmentos terminales obtusos en el ápice y a menudo escasamente emarginados; esporas pardo oscuro; 2n=58, 116?. Bordes de caminos, riberas escarpadas, rocas húmedas. Ch (Breedlove 40252, MO); G (Molina R. et al. 30189, MO); ES (Lellinger, 1989: 126); N (Lellinger, 1989: 126); CR (Jiménez 356, CR). (600-)1900-2900 m. (S. México, Mesoamérica, Venezuela, Ecuador, Perú, Brasil, Eurasia, África, Nueva Zelanda.)

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Rhizomes small, short; scales few, pellucid. Fronds tufted; stipe chestnut-brown, 2-6 cm × 0.5-1 mm, glabrous or occasionally with 1 or 2 pellucid and jointed long hairs; lamina yellowish green, ovate-deltoid to ovate-lanceolate, 4-8 × 1.5-3 cm, submembranous when dry, glabrous, 2-pinnate to 2-pinnate-pinnatifid. Basal pinnae usually larger than adjacent ones; distal pinnae gradually reduced. Ultimate pinnules or segments obovate or obdeltoid, base cuneate and decurrent, apex shallowly lobed. Veins visible on both surfaces. 2n = 58.

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Anogramma leptophylla (L.) Link, Fil. Spec. 137. 1841; Polypodium leptophyllum L. Fig. 13 A, B.

Hojas 2–30 cm de largo y ca 5 cm de ancho, con tendencia al dimor­fismo, anuales; pecíolo pajizo a café-rojizo en la base; lámina lanceo­lada, 2- ó 3-pinnada; últimos segmentos obova­dos, base cunea­da, ápice obtu­so y a menudo escasamente emargina­do.

En Nicaragua esta especie se podría encontrar en bosques de pinos y bosques secos; México a Perú y Brasil, Eurasia, áfrica y Nueva Zelanda.

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Rhizome short-creeping, 1 mm diam., with 1 mm long hairs; fronds 2-25 cm long, clumped; stipe ½-2/3 the frond length, stramineous to reddish-brown at base, glabrous except for scattered hairs 1 mm long at base; blade lanceolate, bipinnate to tripinnate; pinnae 4-6 pairs, segments obovate, cuneate at base, apex obtuse, glabrous; sori naked, along veins; spores dark brown.

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Rhizomes short-creeping, 1 mm diam., with hairs 1 mm long; fronds 2–25 cm long, clumped; stipes 1/2–2/3 the frond length, stramineous to reddish brown at bases, glabrous except for scattered hairs 1 mm long at bases; blades lanceolate, 2–3-pinnate; pinnae 4–6 pairs; segments obovate, cuneate at bases, apices obtuse, glabrous; spores dark brown; 2n=54 (Nay), 58 (Oax, NZ), ca. 116 (India).

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Pflanze einjährig, schon im Juni absterbend. Blattspreite 3–10 cm lang, dünn und zart, unregelmässig 1–3fach gefiedert, die grösseren Abschnitte letzter Ordnung fächerig geteilt, am Grund keilförmig verschmälert. Sori länglich bis lineal, ra

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Rhizome small, set with small brown scales. Fronds few, tufted, erect. Lamina up to 70 x 28 mm, 2- or 3-pinnatifid. Pinnae ovate-deltate. Stipe castaneous, up to 80 mm long. Sori borne along the forked veins.

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Terrestrial annual. Rhizome minute. Fronds few, tufted, erect to slightly arching, 2- or 3-pinnatifid, lamina ovate to oblong, up to 50 x 30 mm, fertile fronds slightly longer and more erect than sterile fronds, glabrous. Sporangia in exindusiate sori borne clustered along veins in lobes of pinnae.

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İnce yapılı, 15 cm’e kadar uzayabilen biryıllık, kısa rizomlu bitkiler. Gametofit çokyıllık, böbreksi, iki loplu, 7-10 mm. Sporofit yaprağı yeşil-sarı renkli, çıplak; genç yapraklar böbrek şeklinde ve 3-5 loplu, 0,8 × 1 cm; olgun yapraklar 2 teleksi, 5-15 × 1-1,5(- 2) cm; yaprak sapı çıplak, 2,5-7 cm, alt kısımda koyu kahverengi-siyah, üst kısımda sarı-yeşil renkli. Telek az teleksi, çıplak, 0,5-2 × 0,5-1 cm, çatallı belirgin damarlı; telek sapı 1-2 mm. Telekçikler genellikle 2 loplu. Sorus turuncu renkli şerit halinde, teleklerin loplarının alt kısmında çok sayıda bir arada bulunur. İndizyum veya yalancı indizyum bulunmaz.

Annual with minute, erect rhizome. Fronds few, tufted, erect, stipe glabrous, lamina 2- or 3-pinnatifid, to 7 cm long, ultimate segments broadly cuneate, shallowly lobed. Null Sori along veins, exindusiate.

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Rhizome minute, annual, set with minute, pale, linear, entire rhizome-scales c. 1 mm long. Fronds few, tufted erect, herbaceous; stipe castaneous, up to 80 mm long, glabrous except for a few small pale scales basally; lamina oblong-ovate to narrowly deltate, up to 70 x 28 mm, 2- to 3-pinnatifid (rarely pinnate in very small plants); pinnae ovate-deltate; ultimate segments broadly to narrowly cuneate, emarginate or shallowly lobed, glabrous on both surfaces, bearing the sori; rhachis castaneous to stramineous, glabrous, narrowly winged in upper part.

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3–4. Feuchte, schattige Felsnischen und Grotten, auf kalkarmem Gestein, kollin-montan. VS, TI.

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Base of quartzite cliffs in moist situations.

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Mesic Grassland. Damp, shady, moist earth slopes.

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Damp, shady, moist earth slopes, 150-1000 m.

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Shaded earth banks, c. 150-2600 m.

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Terrestrial on mossy banks in oak and other deciduous forests; Etla, Ixtlán, Juquila, Miahuatlán, Pochutla, Putla, Sola de Vega, Teo-titlán, Tlaxiaco, Villa Alta; 800-2600 m. Mexico (Chih, Sin, Jal, Col, Mich, Gro, Mex, Mor, Ver, Tam, Oax, Chis); Guat to Pan; Ec, Peru; Old World.

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Terrestrial on mossy banks in oak and other deciduous forests; 800–2800 m. Mexico; Guat, Salv, Nic, CR, Pan; Ven, Ec, Peru, Braz; Eurasia, Africa, NZ.

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Gifberg to Worcester, nearly worldwide.

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Cape Province, Natal and Zimbabwe, Kenya, Uganda, Tanzania, Zaire, Cameroun, Algeria, Sudan and Ethiopia, as well as Canary Islands, Azores, Madeira, S. Europe, Turkey, Nepal, Sri Lanka, Java, Brazil, W. Australia and S. New Zealand.

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