The World Flora Online

8. Adiantum raddianum C. Presl, Tent. Pterid. 158 (1836). Holotipo: Raddi, Pl. Bras. Nov. Gen., t. 78, f. 2 (1825).

Por R.C. Moran.

Adiantum cuneatum Langsd. et Fisch., A. werckleanum H. Christ.

Rizoma cortamente rastrero, compacto, las escamas 2-4 x 0.5-1 mm, linear-lanceoladas, enteras, concoloras, castaño; pecíolo glabro; lámina 10-30 x 6-18 cm, deltada u ovada, 2-4-pinnada, glabra en ambas superficies, verde (no glauco); pinnas 6-10 pares, la más grande 5-20 x 3-15 cm, conspicuamente pediculada, la pínnula acroscópica basal pinnada y sin traslaparse al raquis; raquis y costas glabros, sin una mancha blanca en la axila; últimos segmentos 7-15 x 6-15 mm, flabelados a obovados, a menudo simétricos, lobados o profundamente incisos, pediculados, el pedículo 2-5 mm, no dilatado o articulado en el ápice, el color oscuro pasando a la base del segmento y desvaneciéndose gradualmente; nervaduras de los segmentos estériles terminando en senos; soros 2-9 por segmento, circulares, sin farina amarilla entre los esporangios. Acantilados, bordos sombreados. Ch (Breedlove 27375, MO); G (Croat 41145, MO); N (Stevens 17648, MO); CR (Scamman, 1960: 17). 200-2300 m. (S. México, Mesoamérica, Colombia, Ecuador, Perú, Bolivia, Brasil, Paraguay, Uruguay, Chile, Antillas, Trinidad.)

Source: [
• 3
• 4
]. 

Adiantum raddianum C. Presl, Tent. Pterid. 158. 1836. Fig. 4 E.

Rizoma cortamente rastrero, con escamas linear-lanceoladas, cafés, enteras; hojas hasta 60 cm de largo; pecíolo glabro; lámina ovado-deltada a ovado-lanceolada, 2–4-pinnada, glabra; raquis glabro, algo flexuoso en su 1/2 distal; pinnas 6–10 pares; últimos segmentos obovados a romboidales, márge­nes dista­les variadamente lobulados, bases ampliamente cuneadas a redondeadas, pedículos no dilata­dos en el ápice, no articulados, el color oscuro del pedículo pasando al segmento y desvaneciéndose gradual­mente; en los segmentos estériles los nervios termi­nando en los senos de los márgenes dentados; soros 2–9 por segmento, indusios reniformes a reniforme-redon­­deados sobre el margen distal del seg­mento.

Bosques húmedos; Stevens 17648; 550 m; México a Chile y Brasil, las Antillas, y en el Viejo Mundo. EN

Source: [
• 5
]. 

Rhizome compact; rhizome scales dark brown, linear-lanceolate, entire, 2.5-3.0 mm long, 0.5 mm wide; fronds clumped, arching, ca. 15-45 cm long; stipe ca. ½ the frond length, dark reddish-brown, glabrous except for few scales at base; blade ovate-deltate to ovate-lanceolate, 2-3 times pinnate; pinnae 6-9 pairs, tapering to apex, pinnae and pinnules stalked (1-5 mm long); segments not articulate, the stalk color running into segment base, obovate to rhomboid, 8-12 mm long, sterile margin dentate with veins ending in sinuses, upper and lower surfaces glabrous; sori many per segment; indusium reniform to orbicular, 1-1.5 mm long, glabrous; spores tan.

Source: [
• 7
]. 

Rhizomes short-creeping, 3–5 mm diam.; rhizome scales dark to light brown, linear-lanceolate, entire, 2.5–3 x 0.5 mm; fronds clumped, ca. 20–45 cm long, laxly arching; stipes dark reddish brown, 10–20 cm x 1–2 mm, ca. 1/2 the frond length, glabrous except at bases; blades ovate-deltate to ovate-lanceolate, 2–3- pinnate proximally, 1–2-pinnate distally, tapering to apices; rachises red-brown, glabrous; pinnae 6–9 pairs, pinnae and pinnules with stalks 1–5 mm long; pinnulets obovate to rhomboid, 8–12 mm long, sterile margins dentate, the stalk color passing into pinnulet bases, non-articulate; veins free, forking, ending in sinuses; indument absent on both surfaces; idioblasts absent; sori 4–6 per pinnulet, confined to distal margins; indusia 1–1.5 mm long, reniform to round-reniform, glabrous.

Source: [
• 8
]. 

Tallo: rizoma(s) corta(s)/rastrera(s)/suberecto(s)/nodoso(s). Hoja: fronde(s) 2 pinada(s)/3 pinada(s)/4 pinada(s); raquis glabro(s); estéril(es) margen(es) denticulada(s); nervura(s) libre(s); idioblasto inconspicuo(s); pina o pínula(s) contínua(s)/corta(s)/peciolulada(s)/no dimidiado/glabro(s)/en la(s) amba(s) superficie(s). Tipo de esporangio: leptosporángio pedicelado(s)/glabro(s). Esporangio: ánulo vertical/con/indusio/glabro(s). Esporo: trilete arrugada(s)/à/tuberculada(s).

Source: [
• 1
]. 

Caule: rizoma(s) curta(s)/rasteira(s)/subereto(s)/nodoso(s). Folha: fronde(s) 2 pinada(s)/3 pinada(s)/4 pinada(s); raque glabro(s); estéril(eis) margem(ns) denticulada(s); nervura(s) livre(s); idioblasto inconspícuo(s); pina ou pínula(s) contínua(s)/curta(s)/peciolulada(s)/não dimidiado/glabro(s)/na(s) amba(s) superfície(s). Tipo de esporângio: leptosporângio pedicelado(s)/glabro(s). Esporângio: ânulo vertical/com/indúsio/glabro(s). Esporo: trilete rugada(s)/à/tuberculada(s).

Source: [
• 1
]. 

Stem: rhizome short/creeping/suberect/nodose. Leaf: frond 2 pinnate/3 pinnate/4 pinnate; rachis glabrous; sterile margin denticulate; veins free; idioblast inconspicuous; pinna or pinnule continuous/short/petiolulate/not dimidiate/glabrous/on the both surface. Type of sporangium: leptosporangium pedicellate/glabrous. Sporangium: annulus vertical/with/indusium/glabrous. Spore: trilete rugate/to/tuberculate.

Source: [
• 1
]. 

Rhizome creeping, set with rhizome scales up to 1.5 mm long. Fronds 3- or 4-pinnate. Veins of sterile pinnules ending in sinuses between marginal teeth or crenations of teeth. Ultimate segments not articulated, persistent. Sori orbicular.

Source: [
• 6
]. 

Plants terrestrial or epilithic. Rhizome short-creeping, irregularly branched, to 50 mm long, to 2 mm in diameter, set with roots, closely spaced persistent stipe bases and scales, scales firmly chartaceous, castaneus, adnate, subulate to narrowly triangular, entire to denticulate, apex terminates in a short subulate cell, to 1 mm long, to 0.3 mm wide. Fronds to 4 mm apart, erect to arching, to 640 mm long; stipe firm, atrocastaneus, nitid, terete, to 350 mm long, to 1.2 mm in diameter, basally set with scales similar to those on rhizome, to 2 mm long, to 0.5 mm wide, glabrous apically; lamina anadromous, ovate to broadly ovate, to 3-pinnate, to 300 mm long, to 220 mm wide, with up to 12 pinna pairs; rachis and lower order axes atrocastaneus, nitid, terete, glabrous; pinnae petiolate, petiole to 25 mm long, alternate, basal pinnae largest, more widely spaced than apically, usually overlapping, to 2-pinnate, ovate, to 130 mm long, to 90 mm wide, with up to 8 pinnule pairs; pinnules petiolate, petiole to 8 mm long, alternate, usually overlapping, ovate, to 55 mm long, to 36 mm wide, with up to 5 segment pairs; ultimate segments petiolate, petioles to 2 mm long, alternate, thinly herbaceous, flabellate, narrowly to broadly cuneate, variously lobed, lobes dentate, to 10 mm long, to 10 mm wide, glabrous adaxially and abaxially. Venation evident, flabellately forked, ending in margin in sinus between teeth. Sori borne along outer margin of ultimate segment lobes, on soral flaps which veins enter, soral flaps membranous, subcircular to reniform, to 2 mm in diameter; sporangia confined to veins, sessile, capsule circular to obovate in lateral view, with 19(-21) indurated annulus cells, epistomium (4-)5-celled, hypostomium (4-)5(-6)-celled, stomium with 3 or 4 narrow cells of which walls are conspicuously thickened; Spores brown, tetrahedral-globose, trilete, rugulose, exospore (40-)44(-48) µm in equatorial diameter.

Source: [
• 9
]. 

Perennial with widely spreading rhizome. Fronds arching, stipe glabrous, lamina ovate-deltate, 3- or 4-pinnate, ultimate segments obcuneate, outer margins minutely cuneate-serrate, veins ending in sinuses between marginal teeth. Sori on outer margins of ultimate segments, indusial flaps reniform.

Source: [
• 10
]. 

Erva

Streambanks in forest between 150 and 1310 m.

Source: [
• 11
]. 

Rupícola

Terrestrial or epilithic, usually in disturbed sites such as earthbanks along roads, but often also in undisturbed habitats, usually in light shade. Not edaphically bound.

Source: [
• 9
]. 

Terrícola

Forest, stream banks.

Source: [
• 6
]. 

Naturalized on moist stream banks, 100-250 m.

Source: [
• 10
]. 

Native to South America.

Source: [
• 6
]. 

Terrestrial, on open or lightly shaded banks; Juquila, Putla; 1350-1850 m. Mexico (Tlax, Oax, Chis); Guat to CR; L Antill; Col to Braz & Parag.

Source: [
• 7
]. 

Terrestrial on open or lightly shaded banks, oak forests, seasonal evergreen forests; 800–2200 m. Mexico; Guat, Nic, CR; Gr & L Ant; Col, Ven, Trin, Ec, Peru, Braz, Bol, Parag, Arg, Uru; widely naturalized in OW tropics.

Source: [
• 8
]. 

A native of South America now naturalised in many parts of sub-Saharan Africa.

Source: [
• 9
]. 

S American weed, Cape Peninsula to Wilderness.

Source: [
• 10
]. 

Cape Province, Transkei, Natal, Transvaal, Zimbabwe, Mozambique, Malawi, Tanzania, São Tomé\St Helena, Mauritius and Ascension Island.

Source: [
• 11
].