Stems with hyalodermis present or absent. Leaves radially and spirally arranged, lanceolate, oblong-lanceolate, linear-lanceolate, or oblong-elliptic, symmetric, weakly concave to keeled; base broad, decurrent, tapered, scarcely tapered, or slightly rounded to insertion; margins often irregularly serrulate at apex; laminal cell abaxial and adaxial surfaces plane to somewhat domed, usually 1-papillose, papillae prominent, walls collenchymatous or not. Specialized asexual reproduction by leaflike propagula or absent. Sexual condition dioicous; perigonia as terminal discoid splash platforms, or budlike in series down stem; perichaetial paraphyses absent or of 9-12 cells. Seta usually single, red basally, yellow distally. Spores 8-20 µm, smooth or slightly papillose.