Plants large, in loose tufts, light green or whitish. Stems creeping, densely to moderately densely terete-foliate, usually julaceous, regularly pinnate, branches terete-foliate; central strand present; pseudoparaphyllia acute; axillary hairs of 3-5 cells. Stem leaves erect, loosely imbricate to somewhat spaced proximally, closely imbricate near shoot apices, ovate or elliptic proximally, strongly concave, not or slightly plicate; base broadly long-decurrent; margins serrate to serrulate; apex abruptly tapered, piliferous; costa to 30-60% leaf length, moderate to somewhat stout, terminal spine absent; alar cells subquadrate, enlarged, ; laminal cells linear, walls thick; basal juxtacostal cells shorter, sometimes broader. Branch leaves similar, smaller, narrower, or often more gradually tapered and twisted just below apex. Sexual condition dioicous; perichaetial leaves abruptly contracted to long, reflexed acumen. Seta dark brown, rough. Capsule inclined to horizontal, brown, ovate to elongate, slightly curved; annulus separating by fragments; operculum long-rostrate; peristome xerocastique, perfect. Calyptra naked. Spores 12-15 µm.