Crassula L.
  • Sp. Pl. 1: 282. 1753. (1 May 1753)
  • Pygmyweed [Latin crassus, thick, and -ula, diminutive, alluding to leaves]


Cite taxon page as 'WFO (2021): Crassula L. Published on the Internet;http://www.worldfloraonline.org/taxon/wfo-4000009587. Accessed on: 28 Sep 2021'

General Information

1. Crassula L.

Tillaea L.

Por F.R. Barrie.

Hierbas perennes o subarbustos, rara vez hierbas anuales; tallos típicamente suculentos, en Mesoamérica, cuando hierbas acuáticas o subacuáticas anuales, entonces glabras con tallos delgados no suculentos. Hojas opuestas, simples, las bases connatas y más o menos envainadoras; pecíolo ausente. Inflorescencias axilares, cimosas, las flores 2 o 3 o solitarias; brácteas 2, connatas basalmente. Flores sésiles, subsésiles o pediceladas, los pedicelos 1-4 mm; el cáliz de 3-5 sépalos, connatos basalmente; corola de 3-5 pétalos, libres; estambres 3-5; escamas del receptáculo 0.5-0.8 mm, filamentosas; carpelos 3-5, libres; óvulos 1-numerosos. Semillas 1-numerosas, las superficies lisas, estriadas y/o papilosas. Aprox. 200 spp. Continente americano, Europa, Asia, pero en su mayoría de Sudáfrica; 3 spp. en Mesoamérica.

Las aprox. 15 spp. americanas pertenecen a un grupo de hierbas anuales efímeras creciendo en ciénagas, acuáticas y subacuáticas, antes formalmente segregadas bajo el nombre de Tillaea. Sin embargo, estudios moleculares indican que el grupo está anidado dentro de Crassula (Thiede y Eggli, 2006). Las especies tienen extensas áreas de distribución pero solo han sido colectadas esporádicamente.

Bibliografía: Bywater, M. y Wickens, G.E. Kew Bull. 39: 699-728 (1984).

  • Provided by: [G].Flora Mesoamericana
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    Herbs [shrubs], annual or perennial, aquatic or terrestrial, not viviparous, 0.1-5 dm, glabrous [pubescent]. Stems erect, decumbent, or spreading, simple or branching, succulent. Leaves persistent or deciduous, cauline, opposite, sessile, connate basally; blade ovate, oblong, triangular to lanceolate or oblanceolate, or linear, laminar, 0.1-7 cm, fleshy, base not spurred, margins entire, with glands (hydathodes) in submarginal rows [scattered]; veins not conspicuous. Inflorescences thyrses or panicles [solitary flowers] in axils of leaves (flowers clustered when distal leaves smaller and crowded). Pedicels present. Flowers erect, 3-4(-5)-merous; sepals connate basally, all alike; petals spreading or recurved, distinct [connate], whitish; calyx and corolla not circumscissile in fruit; nectaries linear [various]; stamens as many as sepals; filaments free; pistils spreading to erect, distinct; ovary base rounded; styles 2+ times shorter than ovary. Fruits slightly recurved or ascending to erect. Seeds oblong or ellipsoid to reniform, ridged, sometimes also papillate. x = 8 (secondarily 7).

  • Provided by: [D].Flora of North America @ efloras.org
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    "Fls (3)4–5(–9)-merous; pet distinct or united only at base; stamens as many as and alternate with the pet, adnate to the base of the sep; carpels distinct or nearly so, the style short or even wanting, the stigma terminal; fr follicular; seeds (1–) several or many; succulent herbs or shrubs with thick, opposite lvs and mostly small fls. 250, cosmop."

  • Provided by: [F].New York Botanical Garden
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    Morphology

    Stamens equal in number to the petals, episepalous, or in gamopetalous corollas affixed to the tube; filaments often subulate, sometimes flattened; anthers ovate or oblong, sometimes with conspicuous connective

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    Nectary scales usually very small, narrowly to broadly spathulate to obcuneate or square, rounded and slightly emarginate at the apex, rarely stipitate

  • Provided by: [E].Plants Of the World Online Portal - FTEA
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    • 8
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    Carpels free or slightly united at the base; ovules numerous, rarely 1–2; styles often subulate, narrowly cylindrical or filamentous, often as long as the ovaries, or short and relatively thick; stigmata small, terminal or more rarely subdorsal.

  • Provided by: [E].Plants Of the World Online Portal - FTEA
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    • 8
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    Annual or perennial herbs, rarely shrubs, frequently fleshy

  • Provided by: [E].Plants Of the World Online Portal - FTEA
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    • 8
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    Petals usually slightly united at the base, white or red, rarely yellow or bluish; lobes stellate, erect, patent or connivent, apex often dorsally mucronate, the mucro small, hemispherical or cylindrical, usually blunt or conspicuous, ovoid or subglobular

  • Provided by: [E].Plants Of the World Online Portal - FTEA
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    • 8
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    Sepals free or slightly united at the base

  • Provided by: [E].Plants Of the World Online Portal - FTEA
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    • 8
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    Calyx usually much shorter than the corolla

  • Provided by: [E].Plants Of the World Online Portal - FTEA
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    • 8
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    Flowers isomerous, 3–4- or 5-merous, rarely 6- or 9-merous

  • Provided by: [E].Plants Of the World Online Portal - FTEA
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    • 8
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    Inflorescence usually corymbose, subumbellate, capitate or thyrsoid, sometimes an axillary fascicle or flowers rarely solitary

  • Provided by: [E].Plants Of the World Online Portal - FTEA
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    • 8
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    Leaves very rarely absent, decussate, free or ± connate at the base to form a sheath around the stem, usually sessile, almost always entire, flattish, semiterete or subterete, glabrous, pubescent papillose or scabrid, margin smooth, ciliate or papillate

  • Provided by: [E].Plants Of the World Online Portal - FTEA
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    • 8
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    Scales shorter than the carpels, hyaline or reddish-brown or rose, thin, narrowly to broadly spathulate or obovate or cuneate.

  • Provided by: [C].Plants Of the World Online Portal - FZ
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    Carpels free or connate at the base, oblong or obovoid, attenuate towards or contracted into a ± short usually terminal style, sometimes the styles nearly absent and stigmas subdorsal, completely glabrous or papillose along the suture; ovules numerous or sometimes 1-4.

  • Provided by: [C].Plants Of the World Online Portal - FZ
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    Stamens free or with the lower part of the filaments connate with the corolla-tube, alternipetalous; anthers ovate or oblong, sometimes nearly circular.

  • Provided by: [C].Plants Of the World Online Portal - FZ
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    Corolla usually white or whitish turning orange or brownish-red when dry, sometimes red or carmine, rarely bright yellow, persistent; petals erect or stellate, connate at the base into a ± short tube.

  • Provided by: [C].Plants Of the World Online Portal - FZ
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    Flowers (3-4)5(6-9)-merous, isomerous, usually small and not showy, in cymes arranged either in dense subsessile or pedunculate axillary clusters sometimes forming thyrsoid inflorescences, or in corymb-like axillary or terminal loose or ± dense inflorescences, sometimes 1(2) flowers in the leaf-axils.

  • Provided by: [C].Plants Of the World Online Portal - FZ
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    Calyx usually shorter than the corolla, with the sepals free or slightly connate at the base, appressed to the corolla, ± succulent.

  • Provided by: [C].Plants Of the World Online Portal - FZ
    • Source: [
    • 9
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    Annual or perennial succulent herbs, sometimes with a tuber-like root, or undershrubs or shrubs with ± woody root-stock and succulent usually ± fleshy leaves.

  • Provided by: [C].Plants Of the World Online Portal - FZ
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    Leaves opposite, usually decussate, the lowermost frequently rosulate, free or ± connate in a sheath, usually simple, undivided and entire, thin to ± thick, flat, terete, semiterete, ovoid, etc.

  • Provided by: [C].Plants Of the World Online Portal - FZ
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    Plantes'herbacées, annuelles ou vivaces, à tiges charnues ou plantes semi-arbustives.'Tiges'± ligneuses; racines fréquemment tubéreuses.'Feuilles'décussées, libres ou ± connées à la base, ordinairement sessiles, presque toujours entières, planes, semi-cylindriques ou cylindriques, glabres, papilleuses ou scabres, à marge lisse, ciliée ou papilleuse.'Fleurs ordinairement disposées en inflorescences corymbeuses subombellées, capitées ou thyrsoïdes, rarement solitaires, isomères, haplostémones, le plus souvent 5-mères, rarement 3-4-mères, plus rarement 6-9-mères; calice en général plus court que la corolle, à sépales libres ou légèrement soudés à la base; pétales presque toujours soudés à la base, étalés ou érigés, récurvés ou confluents au sommet, le plus souvent mucronulés ou pourvus d'un appendice épaissi ± globuleux ou capité; étamines épisépales à filet inséré sur le court tube corollin (chez les espèces gamopétales) filiformes ou légèrement cunéées; anthère sovales ou oblongues; carpelles libres ou un peu soudés à la base, rétrécis en un style subulé aussi long ou plus court que l'ovaire; ovules nombreux, rarement 1-2; squamules nectarifères, plus petites que l'ovaire, obcu-néées, arrondies et légèrement émarginées.\n\t\t\t\tCe genre renferme environ 300 espèces, la plupart (230) réparties en Afrique du Sud, les autres en Afrique orientale, en Arabie, à Mada-gascar, en Europe et en Amérique.\n\t\t\t\tLa révision systématique de la section des'Tillaeoideae infirmera ou confirmera les larges répartitions de certaines espèces.

  • Provided by: [H].Flore d'Afrique Centrale
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    Literature

    SELECTED REFERENCES Bywater, M. and G. E. Wickens. 1984. New World species of the genus Crassula. Kew Bull. 39: 699-728. Merxmüller, H., H. C. Friedrich, and J. Grau. 1971. Cytotaxonomische Untersuchungen zur Gattungsstruktur von Crassula. Ann. Naturhist. Mus. Wien 75: 111-119. Moran, R. V. 1992b. Pygmyweed (Crassula connata) etc. in western North America. Cact. Succ. J. (Los Angeles) 64: 223-231. Toelken, H. R. 1977. A revision of the genus Crassula in southern Africa. Contr. Bolus Herb. 8. Van Jaarsveld, E. J. 2003. Crassula. In: U. Eggli, ed. 2003. Illustrated Handbook of Succulent Plants. Crassulaceae. New York. Pp. 32-84.

  • Provided by: [D].Flora of North America @ efloras.org
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    Included Species

    Synonyms

    Other Local Names

    NameLanguageCountry
    Pygmyweed [Latin crassus, thick, and -ula, diminutive, alluding to leaves]

     Information From

    Crassulaceae
    World Flora Online Data. 2017.
    • A CC0 1.0 Universal (CC0 1.0).
    MBG Floras Images
    http://www.tropicos.org/ImageSearch.aspx
    Flora images. Missouri Botanical Garden. Accessed on Jun. 2018.
    • B Missouri Botanical Garden
    Plants Of the World Online Portal - FZ
    https://www.kew.org/science/who-we-are-and-what-we-do/strategic-outputs-2020/plants-of-the-world-online
    http://www.plantsoftheworldonline.org/terms-and-conditions
    • C
    Flora of North America @ efloras.org
    http://www.efloras.org/flora_page.aspx?flora_id=1
    'Flora of North America @ eFloras (2008). Published on the Internet http://www.efloras.org/flora_page.aspx?flora_id=1 [accessed August 2016]' Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
    • D Flora of North America Association
    Plants Of the World Online Portal - FTEA
    https://www.kew.org/science/who-we-are-and-what-we-do/strategic-outputs-2020/plants-of-the-world-online
    http://www.plantsoftheworldonline.org/terms-and-conditions
    • E
    New York Botanical Garden
    Descriptions of plants should be attributed to the full citation for each individual article, chapter or book that is the source for each record, which should include the authors of original publication.
    • F Content licensed under Creative Commons Attribution-NonCommercial-ShareAlike 3.0 Unported License
    Flora Mesoamericana
    http://www.mobot.org/MOBOT/fm/
    Gerrit Davidse, Mario Sousa Sánchez, A. O. Chater, Universidad Nacional Autónoma de México. Instituto de Biología, Missouri Botanical Garden, Natural History Museum (London, England) UNAM, 1994
    • G Missouri Botanical Garden
    Flore d'Afrique Centrale
    https://www.floredafriquecentrale.be
    • H http://creativecommons.org/licenses/by/4.0/ Attribution 4.0 International (CC BY 4.0)
    World Flora Online consortium
    http://www.worldfloraonline.org/organisation/WFO
    World Flora Online Data. 2018.
    • I CC0 1.0 Universal (CC0 1.0).