Herbs, without stinging hairs. Stems ascending. Leaves alternate or opposite; stipules persistent, lateral, free; leaf blade ovate, 3-veined or triplinerved, margin serrate; cystoliths punctate. Inflorescences axillary, androgynous (plants monoecious), sessile, glomerules crowded on broad, campanulate, dentate involucre. Male flowers numerous on margin of the involucre; perianth lobe 1, often split and unequally 3-lobed or 3-toothed; stamen 1; rudimentary ovary absent. Female flowers 1 or 2, in center of involucre; perianth absent; staminodes absent. Ovary straight; stigma filiform, papillose-hispid, almost persistent; ovule orthotropous. Achene compressed ovoid, woolly or glabrous, enclosed by involucre. Seeds with endosperm; cotyledons elliptic.
Flowers in sessile inflorescences, single or few–many together, surrounded by an involucre of fused bracts; inflorescences bisexual or unisexual by abortion (and then usually ♀), axillary and/or in terminal spikes (if each involucre contains only 1(–2) ♀ flowers, the number of involucres in each leaf-axil is usually high)
Male flowers ± boat-shaped; perianth cylindrical to conical at the base, open along one side and with a ± erect tip, enclosing 1 at first inflexed, later reflexed stamen; rudimentary ovary absent
Female flowers without perianth, consisting of a pistil only; style filiform
Achenes enclosed in the persistent involucre.7 species, distributed in tropical and South Africa, Yemen, S. India and Java.
Annual or perennial herbs, occasionally subshrubs, with erect or prostrate stems, usually monoecious (or plants with only ♀ flowers by abortion of all bisexual inflorescences)
Leaves opposite or alternate, petiolate; cystoliths punctiform; stipules lateral, free
Inflorescences of sessile clusters in the leaf axils; clusters either bisexual with few-many flowers surrounded by a common campanulate involucrum of fused bracts or inflorescence clusters of many very small involucres each with 1(2) female flowers (the involucres then appear as perianths).
Bisexual involucres usually with many male flowers and 1 -few female flowers (occasionally all the flowers are male); bracts and flowers with long woolly hairs.
Male flowers pedicellate, club-shaped; perianth 3-lobed, one lobe apiculate and provided with prominent bristles; stamen 1, inflexed in bud, later reflexed; rudimentary ovary absent.
Female flowers naked, pedicellate, consisting of an ovary covered with a woolly tomentum; stigma sessile, filiform; staminodes absent Entirely-female involucres are common on certain individuals, and then much smaller and with much fewer flowers than the bisexual involucres.
Slender annual or perennial herbs, occasionally subshrubs, with erect or prostrate stems, monoecious or apparently dioecious (by abortion of all bisexual inflorescences leaving only female flowers).
Leaves alternate or opposite, petiolate, serrate; stipules lateral, free.
Achene enclosed in the persistent involucre, brown, shiny, often crowned by the persistent stigma.
Herbes'vivaces ou sous-arbustes, monoïques ou dioïques.'Feuilles'alternes ou opposées, pétiolées, serretées ou crénelées, trinerves, à cystolithes punctiformes; stipules libres, latérales.'Inflorescences'en glomérules axillaires, constituées soit par 2 capitules androgynes pluriflores à involucre cupuliforme ou campanulé, denté sur le bord, et de quelques involucres ♀ 1-2-flores, urcéolés, à orifice étroit, dentelé laissant passer les stigmates, soit par de nombreux involucres ♀.'Fleurs'♂ à périgone en forme de cornet 3-lobé couvert de longs poils, à 1 étamine; fleur ♀ apérianthée, à stigmate filiforme.'Akènes nus ou enveloppés par l'involucre ♀ accrescent.\n\t\t\t\tCinq espèces d'Afrique et d'Indo-Malaisie, dont 2 au Congo belge, dans les montagnes de l'Est.