Gouania Jacq.
  • Select. Stirp. Amer. Hist. 263–264, pl. 179, f. 40. 1763. (Jan 1763)
  • Chewstick, jaboncillo [For Antoine Gouan, 1733–1821, French botanist and ichthyologist at Montpellier, director of botanical garden in 1767, later professor of botany and medicine]


Cite taxon page as 'WFO (2021): Gouania Jacq. Published on the Internet;http://www.worldfloraonline.org/taxon/wfo-4000016086. Accessed on: 16 Oct 2021'

General Information

6. Gouania Jacq. N.v.: Chew-stick, B; jaboncillo, G, ES; limpia-dientes, H.

Por A.Pool.

Bejucoso arbustos desparramados, trepadores por zarcillos circinados, los zarcillosoriginados en las axilas de las hojas, especialmente aquellas subyacentes a lasinflorescencias, ramas axilares terminales, o en las axilas de las brácteas dela cima; las plantas inermes; escamas de las yemas presentes. Hojas alternas,pinnatinervias, algunas veces con el par de nervaduras laterales más proximaloriginándose con la vena media en la unión del pecíolo y la lámina, pero estasno más fuertes o solo ligeramente más fuertes que las nervaduras laterales másdistales y no tan fuertes como la vena media, las nervaduras laterales sinfranjas, las glándulas en la unión de la lámina con el pecíolo con frecuenciaestipitadas o foliáceas, los márgenes dentados, cada diente con una glándulaapical; pecioladas; estípulas libres, dispuestas lateralmente en la base delpecíolo, no lobadas o con más frecuencia 2-lobadas o rara vez 3-lobadas.Inflorescencias axilares o terminales, en tirsos racemiformes o paniculadoscompuestos de cimas pequeñas con numerosas flores, las cimas abrazadas por brácteas,los pedúnculos de las cimas generalmente muy cortos, los pedicelos alargándosea medida que las flores maduran. Flores bisexuales (en Mesoamérica; en otraspartes algunas veces unisexuales por aborto, las plantas polígamas), 5-meras;hipanto levemente obcónico a campanulado, adhiriéndose al ovario; sépalostriangulares, adaxialmente carinados, persistentes; pétalos generalmente máscortos que los sépalos a tan largos como estos, marcadamente cóncavos,cortamente unguiculados, el limbo apicalmente redondeado a bífido, blancos overde-blanquecinos, envolviendo los estambres en la antesis; estambres opuestosa los pétalos, ligeramente más cortos que los pétalos en la madurez aligeramente más largos que estos; disco anchamente anular, casi llenando porcompleto la boca del hipanto excepto el pequeño anillo central, epígino,carnoso, nectarífero, glabro, peloso, o los tricomas restringidos al borde delanillo, con 5 lobos cartáceos de posible origen estaminodial opuestos a lossépalos (rara vez no lobado, no en Mesoamérica), los lobos enteros oapicalmente bilobados o irregularmente lobados; ovario ínfero, 3-locular, elestilo trífido, exerto del anillo discoidal generalmente después de ladehiscencia del estambre, los estigmas 3, pequeños. Frutos en esquizocarpossecos, 3-alados al madurar, separándose septicidamente en 3 mericarpos 2-aladoscada uno de los cuales permanece temporalmente unido apicalmente y suspendidosdesde 2 carpóforos; mericarpos indehiscentes, con 1 semilla. Aprox. 50 spp.,pantropical con la mayoría de las especies en América desde los Estados Unidos[S. Florida] hasta Sudamérica, las especies sudamericanas necesitan unarevisión moderna.

Laforma de los mericarpos es muy útil en la distinción de las especies de Gouania. Los mericarpos en forma de mancuernasson transversalmente oblongos con un seno muy ancho en el ápice y la base delcuerpo del fruto (o rara vez solo en el ápice); el cuerpo mismo del fruto es enforma de la barra angosta de la mancuerna. Las alas del mericarpo están unidasprimariamente solo de los lados del cuerpo fructífero (o rara vez de los ladosy la base). Los mericarpos en forma de mariposa tienen el contorno más o menoscircular a transversalmente oblongo con escotaduras en el ápice y la base delcuerpo del fruto (o rara vez solo en la base); las alas están unidas al ápice yla base del cuerpo del fruto así como también a los lados. Para ayudar adefinir la forma de los merica

  • Provided by: [G].Flora Mesoamericana
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    Shrubs climbing, usually with tendrils, unarmed. Leaves alternate, with caducous stipules, petiolate, pinnativeined or 3-veined from base, margin entire or dentate. Flowers polygamous, in terminal or axillary, cymose racemes or cymose panicles, often with tendrils at lower part of rachis or base. Calyx tube short, coadnate with ovary; sepals 5, ovate-triangular, adaxially medially keeled, dehiscent with longitudinal slits. Petals 5, spatulate. Stamens 5, dorsifixed, enfolded by petals. Disk thick, pentagonous or 5-lobed, glabrous or hairy. Ovary inferior, ± deeply immersed in disk, 3-loculed, with 1 ovule per locule; styles cleft to half or deeply cleft. Capsule subglobose, impressed on both ends, crowned by persistent limb of calyx, 3-winged, with 3 rounded locules split from axil at maturity; locules indehiscent or along inner rib narrowly fissured. Seeds 3, red-brown, shiny, obovoid; endosperm thin.

  • Provided by: [J].Flora Of CHina @ efloras.org
    • Source: [
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    • ]. 

    GOUANIA Jacq.

    Lianas or scandent shrubs, climbing by circinate tendrils, the tendrils originating from axils of leaves, especially those subtending inflorescences, terminating axillary branches, or axillary from cyme bracts, plants unarmed, bud scales present. Leaves alternate, petiolate, venation pinnate, lateral veins without stripes, glands at junction of leaf blade and petiole frequently stipitate or foliaceous, margin dentate, each tooth with an apical gland; stipules free, borne laterally at base of petiole, un-lobed or more frequently 2- (or rarely 3-) lobed. Inflorescences axillary or terminal, racemiform or paniculate thyrses composed of small, multi-flowered cymes, the cymes subtended by bracts, peduncles of cymes usually very short, pedicels elongating as flowers mature. Flowers bisexual (in our area; reportedly elsewhere sometimes unisexual by abortion, the plants polygamous), 5-merous. Hypanthium shallow obconical to campanulate, adhering to the ovary. Sepals triangular, adaxially keeled, persistent. Petals usually smaller or equal to sepals in length, strongly concave, short-clawed, limb apically rounded to 2-fid, white or greenish-white, enfolding the stamens at anthesis. Stamens opposite petals, slightly shorter to slightly longer than petals at maturity. . Disk broadly annular nearly filling mouth of hypanthium excluding small central annulus, epigynous, fleshy, nectiferous, glabrous, pubescent, or trichomes restricted to rim of annulus, margin with 5 chartaceous lobes opposite the sepals (rarely un-lobed), the lobes entire or apically 2- or irregularly- lobed. Ovary inferior, 3-locular, style 3-fid, the style exerted through disc annulus, usually after stamen dehiscence, stigmas 3, small. Fruit a dry schizocarp, 3-winged at maturity, separating septicidally into three 2-winged mericarps each of which remains temporarily apically attached and suspended from 2 carpophores. Mericarps indehiscent, 1-seeded. Aprox. 50 species, pantropical with majority of species in the New World from Southern Florida to South America, the South American species in need of modern revision.The shape of the mericarps is very helpful in distinguishing the species of Gouania. The dumbbell-shape mericarps are transversely oblong with a very broad sinus at the apex and base of the fruit body. The fruit body itself is like the narrow bar or handhold of the dumbbell. Bornstein (1989: 165) described this fruit shape as “Fruit wing attached to side of fruit body only,” and I borrow his description. It’s not technically true as a thin margin of the wing continues along the apex and base of the fruit body, but it does help to explain the shape. The other shape I term the butterfly. The butterfly-shape mericarps are more or less circular to transversely oblong with emarginations to deep clefts at the apex and base of the fruit body, or only at the base in G. velutina. Bornstein (1989:165) described this type as “Fruit wing attached to side and base, and/or apex of fruit body” and I borrow again from him. To help define the shape of mericarps, the following measurements were made: height of fruit body, height of mericarp wings, distance between highest points of two wings, width of mericarp, and width of fruit body.

    Un grupo tropical y subtropical con quizás 50 especies; 4 especies se encuentran en Nicaragua and 1 expected. Modified AP 18 Aug 2013

  • Provided by: [I].Flora de Nicaragua
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    Lianas, or scandent to arching shrubs, climbing by tendrils. Leaves alternate, pinnately-nerved or sometimes 3-nerved from the base, mostly dentate, the teeth frequently glandular; petiolate, the stipules narrow and deciduous or sometimes broad and persistent. Inflorescences spicate thyrses composed of glomerules, axillary or terminal. Flowers bisexual or 'polygamous; floral tube obconic to siibcampanulate; sepals 5, persistent; petals 5, white to green white, short-clawed; stamens 5, hidden by the petals; nectariferous disc 5-lobed or pentagonal, the lobes opposite the sepals; ovary inferior, immersed in the disc, 3-locular, the style 3-branched more or less basally, the stigmas minute. Fruit a schizocarp, 3-locular, generally 3-winged, splitting longitudinally along the margin of each wing into 3, 2-winged mericarps; seeds 3, the seed coat shiny.

  • Provided by: [E].Flora de Panama
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    Woody vines [climbing shrubs], tendrils present [rarely absent], unarmed; bud scales present. Leaves deciduous [persistent], alternate; blade not gland-dotted; pinnately veined [3-veined from base]. Inflorescences: axillary or terminal, racemelike or paniclelike thyrses; peduncles and pedicels not fleshy in fruit. Pedicels present or absent. Flowers bisexual [unisexual, plants polygamous]; hypanthium cupulate to campanulate, 1.5–3 mm wide; sepals 5, spreading, pale green, ovate-triangular, keeled adaxially; petals 5 [rarely 0], white to yellowish or greenish, hooded, spatulate, clawed; nectary fleshy, margins 5-lobed [unlobed], lining hypanthium but distally free; stamens 5, enfolded by petals; ovary inferior, 3-locular; styles 3, connate basally. Fruits schizocarps, breaking into 3 2-winged samaras [rarely nutlets].

  • Provided by: [C].Flora of North America @ efloras.org
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    Morphology

    Fruit a dry schizocarpous capsule, late in maturation the endo- and exocarpous tissue near the septa (on both sides of each mericarp) producing 3 broad thin wings (thus each wing is composed of parts of 2 adjacent mericarps; reportedly the wing-production fails in certain Asian and American, but not African, populations, and perhaps the reports are based on observations of immature fruit, for the wing-proliferation is a quite late development).

  • Provided by: [D].Plants Of the World Online Portal - FTEA
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    Cymes usually small, but usually aggregated into ample leafless terminal panicles

  • Provided by: [D].Plants Of the World Online Portal - FTEA
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    Stipules present, free, usually caducous

  • Provided by: [D].Plants Of the World Online Portal - FTEA
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    Leaves alternate, petiolate; blades usually ovate, serrate (rarely nearly entire), acute or acuminate, at base usually rounded or cordate and often 3–5-nerved but penninerved more distally

  • Provided by: [D].Plants Of the World Online Portal - FTEA
    • Source: [
    • 6
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    Climbing shrubs or lianes with circinnate tendrils (rarely, or apparently never in Africa, seeming to be mere shrubs)

  • Provided by: [D].Plants Of the World Online Portal - FTEA
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    Disk rather massive, epigynous

  • Provided by: [D].Plants Of the World Online Portal - FTEA
    • Source: [
    • 6
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    Ovary usually urceolate; style trifid or obscurely 3-lobed

  • Provided by: [D].Plants Of the World Online Portal - FTEA
    • Source: [
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    Flowers 5-merous, bisexual, epigynous at all stages

  • Provided by: [D].Plants Of the World Online Portal - FTEA
    • Source: [
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    Flowers usually bisexual.

  • Provided by: [F].Plants Of the World Online Portal - FZ
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    Inflorescence a paniculate thyrse.

  • Provided by: [F].Plants Of the World Online Portal - FZ
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    Stipules small, caducous.

  • Provided by: [F].Plants Of the World Online Portal - FZ
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    Leaves alternate, petiolate; lamina entire or dentate, penninerved but often 3–5-nerved from the base.

  • Provided by: [F].Plants Of the World Online Portal - FZ
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    Climbing shrubs or lianes; branches provided with coiled tendrils.

  • Provided by: [F].Plants Of the World Online Portal - FZ
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    Sepals 5.

  • Provided by: [F].Plants Of the World Online Portal - FZ
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    Petals 5, cucullate; inserted below the margin of the disk.

  • Provided by: [F].Plants Of the World Online Portal - FZ
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    Stamens 5.

  • Provided by: [F].Plants Of the World Online Portal - FZ
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    Disk filling the receptacle, 5-lobed.

  • Provided by: [F].Plants Of the World Online Portal - FZ
    • Source: [
    • 7
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    Ovary inferior, immersed in the disk; style 3-fid or obscurely 3-lobed.

  • Provided by: [F].Plants Of the World Online Portal - FZ
    • Source: [
    • 7
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    Fruit a schizocarp, longitudinally 3-winged (or 3-angled), septicidal, separating into 3 woody or coriaceous indehiscent cocci.

  • Provided by: [F].Plants Of the World Online Portal - FZ
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    Seeds compressed or planoconvex.

  • Provided by: [F].Plants Of the World Online Portal - FZ
    • Source: [
    • 7
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    Arbustes'le plus souvent sarmenteux ou lianes, s'accrochant en général à l'aide de vrilles circinées.'Feuilles'à stipules caduques, pétiolées, ovales à cordées, entières ou dentelées, à nervation penninerve mais souvent en plus 3-ou 5-nervées à la base.'Fascicules'sessiles à courtement pédonculés, groupés en inflorescences terminales composées spiciformes ou racémeuses.'Fleurs'petites,'☿, moins souvent ♂♀; réceptacle court; calice à 5 dents valvaires; pétales cucullés; disque cachant le fond du réceptacle; ovaire infère; style trifide.'Capsules'à 3 angles souvent ailés, se séparant en 3 coques caduques, indéhiscentes ou déhiscentes par une fente étroite.'Graines comprimées ou planconvexes; albumen peu abondant; radicule courte, infère.\n\t\t\t\tGenre répandu dans presque toutes les régions tropicales, comprenant 66 espèces, 5 en Afrique dont 2 au Congo belge.

  • Provided by: [H].Flore d'Afrique Centrale
    • Source: [
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    Habit

    Lianas shrubs

  • Provided by: [E].Flora de Panama
    • Source: [
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    Distribution

    A genus of 30-50 species, pantropical but mostly New World.

  • Provided by: [E].Flora de Panama
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    Included Species

    Other Local Names

    NameLanguageCountry
    Chewstick, jaboncillo [For Antoine Gouan, 1733–1821, French botanist and ichthyologist at Montpellier, director of botanical garden in 1767, later professor of botany and medicine]

     Information From

    MBG Floras Images
    http://www.tropicos.org/ImageSearch.aspx
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    • A Missouri Botanical Garden
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    http://www.efloras.org/flora_page.aspx?flora_id=1
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    • C Flora of North America Association
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    http://www.mobot.org/MOBOT/fm/
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    • G Missouri Botanical Garden
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    • H http://creativecommons.org/licenses/by/4.0/ Attribution 4.0 International (CC BY 4.0)
    Flora de Nicaragua
    http://www.tropicos.org/projectwebportal.aspx?projectid=7&pagename=Home&langid=66
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    'Flora of China @ eFloras (2008). Published on the Internet http://www.efloras.org/flora_page.aspx?flora_id=2 [accessed August 2016]' Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
    • J Missouri Botanical Garden
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    • L CC0 1.0 Universal (CC0 1.0).