Homalium Jacq.
  • Enum. Syst. Pl. 5, 24. 1760. (Sept-Oct 1760)


Cite taxon page as 'WFO (2021): Homalium Jacq. Published on the Internet;http://www.worldfloraonline.org/taxon/wfo-4000018148. Accessed on: 23 Oct 2021'

General Information

HOMALIUM Jacq.

Por Amy Pool

Homalium racemosum Jacq., Enum. Syst. Pl. 24. 1760; H. nicaraguense S.F. Blake.

Arboles siempreverdes, 5–25 m de alto, inermes; plantas hermafroditas. Hojas alternas, dísticas, elípticas o lanceoladas, 6.5–21 cm de largo y 2.5–7.5 cm de ancho, ápice acuminado, base caudada a redondeada, margen serrado- o undulado-glandular, sin marcas pelúcidas, glándulas basales ausentes, pinnatinervias; pecíolo 0.3–1 cm de largo, estípulas no observadas. Inflorescencias de racimos (a veces ligeramente ramificados) axilares o terminales, 5–9 (–17) cm de largo y 1–2 cm de ancho, con 8–20 flores, pedúnculo 2–3.5 cm de largo, brácteas pequeñas, tempranamente caducas; tubo del cáliz 1–3 mm de largo, lobos (5) 6 ó 7, valvados, 2.5–3.5 mm de largo y 1.2–2 mm de ancho; pétalos 6 ó 7, 3–4.5 mm de largo y 2–2.5 mm de ancho, blancos o verde pálidos; estambres en 6 ó 7 fascículos epipétalos, 2–6 estambres por fascículo, fascículos alternos con las glándulas del disco, glándulas vellosas; ovario semiínfero, parte libre cónica, velloso, estilo 3-fido, estigmas simples. Fruto una cápsula subrómbica, 4–7 mm de largo y 3–5 mm de ancho, tomentosa; semillas 1 ó 2, 2.5 mm de largo, glabras, sin arilo.

Común, áreas alteradas, en todo el país; 5–1200 m; fl dic–may, fr mar–may, sep; Stevens 7440, Vincelli 349; México hasta Perú y Brasil y también en las Antillas. Género pantropical con ca 200 especies, 3 de ellas en América tropical. "Areno".

  • Provided by: [E].Flora de Nicaragua
    • Source: [
    • 9
    • ]. 

    Arbustos o árboles, hermafroditas; estípulas ausentes o diminutas y caducas. Hojas alternas, la lámina usualmente glandular-aserrada o (a veces) glandular-crenada, de otra forma sin glándulas, sin puntos o líneas translúcidos, pinnadamente nervada. Infls. axilares o subterminales, espiciformes o racemosas a paniculadas. Fls. bisexuales, con un disco glandular; sépalos 5–8(–12), connatos basalmente en un hipanto adnato a la parte inferior del ovario, los lóbulos valvados a subimbricados, endurecidos y acrescentes; pétalos 5–8(–12), insertos en la base de la parte superior del ovario, imbricados; estambres numerosos, epipétalos en fascículos de (2)3 ó 4(–6), los filamentos separados, filiformes; ovario semi-ínfero; estilos 2–6(–8), que forman una columna; estigmas simples o capitados. Frs. capsulares, indehiscentes o incompletamente 2–8-valvados desde el ápice; semillas 1–pocas, sin arilo.

  • Provided by: [F].Manual de Plantas de Costa Rica
    • Source: [
    • 11
    • ]. 

    Trees or shrubs. Leaves alternate, rarely opposite or verticillate; stipules caducous; usually petiolate; leaf blade pinnate-veined, margin with glandular teeth, rarely entire. Flowers bisexual, epigynous, small, in terminal or axillary, many flowered racemes or panicles, inserted singly along rachis, or in sessile to shortly pedunculate fascicles; bracts small, caducous or persistent; pedicels slender in flower, articulate at or above middle. Sepals and/or petals often accrescent after anthesis. Calyx tube obconic, adnate to lower 2/3 of ovary and later to lower 2/3 of capsule; sepals (4 or)5-8(-12), spreading, linear, oblong, or obovate-spatulate, persistent. Petals inserted at rim of calyx tube, usually isomerous with and similar to sepals, alternating with them. Disk glands 1 opposite each sepal, rarely more or fewer, small, fleshy, ± globose and hairy. Stamens inserted singly or in groups before each petal, alternating with disk glands and inserted between them, usually finally overtopping perianth; filaments free, filiform; anthers subglobose, small, dorsifixed. Ovary semi-inferior, only upper conic part free above adnate calyx tube, 1-loculed; placentas 2-6(-8), with (1-)3-7 ovules near apex of each placenta; styles 2-5(-7), filiform, free or united in lower 1/3 or less, free parts divergent, usually finally overtopping perianth; stigmas capitate to punctiform, small. Capsule obconic, small, for most of its length enclosed in adnate calyx tube and persistent perianth segments, leathery, apex 2-8-valvate; styles ± persistent. Seeds 1 to few.

  • Provided by: [G].Flora Of CHina @ efloras.org
    • Trees or shrubs. Leaves usually alternate, petiolate, the stipules minute and caducous, rarely absent, the blade entire to usually serrate-crenate, the teeth glandular beneath, penninerved, without pellucid glands. Inflorescences axillary, racemose or paniculate. Flowers , sessile or pedicellate, the pedicels articulated, the bracts usually minute, caducous or + persistent; calyx tube (or receptacle?) turbinate, adnate to the base of the ovary; calyx lobes 5-8(-12), usually narrow, persistent or accrescent; petals isomerous with the calyx lobes, inserted in the throat of the calyx, alternating with the calyx lobes, persistent or accrescent; stamens epipetalous, isomerous with the petals or in fascicles of 2-8(or more), the filaments filiform, the anthers small, extrorse, dorsifixed, longitudinally dehiscent; disk represented by a gland opposite each sepal and alternating with the filaments or fascicles of filaments; ovary with the lower half adnate to the calyx tube (or half inferior?), 1-locular, the placentas parietal, 2-6(-8), each with (1-)2-7 ovules near the apex, the styles isomerous with the placentas, the stigmas simple or capitellate. Fruits capsular, half inferior, ? coriaceous, indehiscent or incompletely 2-8-valvate from the apex; seeds few or solitary, small, angulate, the testa crustace- ous; endosperm copious.

    • Provided by: [I].Flora de Panama
    • Morphology

      Calyx-tube (or receptacle) adnate to the ovary

    • Provided by: [D].Plants Of the World Online Portal - FTEA
      • Source: [
      • 7
      • ]. 

      Flowers bisexual, regular, (4–)5–8(–12)-merous, in axillary or subterminal racemes or panicles, solitary or fasciculate on the inflorescence-branches, sessile or pedicelled, subtended by small caducous or persistent bracts

    • Provided by: [D].Plants Of the World Online Portal - FTEA
      • Source: [
      • 7
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      Petals alternating with the sepals, persistent, sometimes accrescent

    • Provided by: [D].Plants Of the World Online Portal - FTEA
      • Source: [
      • 7
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      Sepals flat, usually narrow, persistent, often accrescent and wing-like

    • Provided by: [D].Plants Of the World Online Portal - FTEA
      • Source: [
      • 7
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      Disk represented by a usually hairy gland opposite each sepal

    • Provided by: [D].Plants Of the World Online Portal - FTEA
      • Source: [
      • 7
      • ]. 

      Stamens epipetalous, solitary or in fascicles of 2–3; filaments slender; anthers small, extrorse, dorsifixed

    • Provided by: [D].Plants Of the World Online Portal - FTEA
      • Source: [
      • 7
      • ]. 

      Capsule half-inferior, coriaceous or woody, finally 2–3(–8)-valved from the apex, or remaining indehiscent (apparently mainly in sterile flowers)

    • Provided by: [D].Plants Of the World Online Portal - FTEA
      • Source: [
      • 7
      • ]. 

      Ovary conical and free in its upper half, sunk in the receptacle in its lower half, 1-locular, with 2–3(–8) placentas, each with 1-several ovules near the apex; styles 2–4(–7), free or connate to a distinct column below (then only stigmatic arms free); stigmas punctiform

    • Provided by: [D].Plants Of the World Online Portal - FTEA
      • Source: [
      • 7
      • ]. 

      Seeds 1–few, small, often not fully formed.

    • Provided by: [D].Plants Of the World Online Portal - FTEA
      • Source: [
      • 7
      • ]. 

      Leaves alternate, simple, entire or glandular-serrate-crenate, penninerved, petiolate; stipules absent, or minute to large, sometimes auriculate, caducous or ± persistent

    • Provided by: [D].Plants Of the World Online Portal - FTEA
      • Source: [
      • 7
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      Trees or shrubs

    • Provided by: [D].Plants Of the World Online Portal - FTEA
      • Source: [
      • 7
      • ]. 

      Trees or shrubs.

    • Provided by: [H].Plants Of the World Online Portal - FZ
      • Source: [
      • 8
      • ]. 

      Flowers bisexual in axillary or terminal racemes or paniculate racemes, solitary or fasciculate on the inflorescence-branches, sessile or pedicelled, subtended by small caducous or persistent bracts.

    • Provided by: [H].Plants Of the World Online Portal - FZ
      • Source: [
      • 8
      • ]. 

      Leaves alternate (in African spp.), petiolate; lamina entire or crenate-serrate with a gland near the apex of each tooth on the underside; stipules absent, minute or large and orbicular, caducous or persistent.

    • Provided by: [H].Plants Of the World Online Portal - FZ
      • Source: [
      • 8
      • ]. 

      Receptacle (or calyx tube?) adnate to the ovary; sepals (4) 5–8 (12), usually narrow, sometimes accrescent.

    • Provided by: [H].Plants Of the World Online Portal - FZ
      • Source: [
      • 8
      • ]. 

      Ovary with its lower half sunk in the receptacle, 1-locular with 2–3 (8) placentas each with (1) 2–3 (7) ovules near the apex; styles 2–4 (7), free or variously connate; stigmas small.

    • Provided by: [H].Plants Of the World Online Portal - FZ
      • Source: [
      • 8
      • ]. 

      Disk represented by a usually tomentose gland opposite each sepal.

    • Provided by: [H].Plants Of the World Online Portal - FZ
      • Source: [
      • 8
      • ]. 

      Stamens epipetalous, solitary or in fascicles of 2–3 (or up to 12 outside our area); filaments slender; anthers small, extrorse, dorsifixed.

    • Provided by: [H].Plants Of the World Online Portal - FZ
      • Source: [
      • 8
      • ]. 

      Petals alternating with the sepals, persistent, sometimes accrescent.

    • Provided by: [H].Plants Of the World Online Portal - FZ
      • Source: [
      • 8
      • ]. 

      Capsule half inferior, woody or coriaceous, 2–3 (8) valved from the apex or indehiscent.

    • Provided by: [H].Plants Of the World Online Portal - FZ
      • Source: [
      • 8
      • ]. 

      Seeds solitary or few.

    • Provided by: [H].Plants Of the World Online Portal - FZ
      • Source: [
      • 8
      • ]. 

      Arbrisseaux ou arbres, parfois de grande taille, inermes; écorce se détachant souvent par plaques irrégulières (comme chez les platanes) ; racines tabulaires.'Feuilles'alternes, persistantes, le plus souvent dentées ou crénelées, penninerves; stipules variables, parfois foliacées, réniformes et persistantes. Racèmes, panicules ou thyrses, axillaires ou terminaux, portant des fleurs solitaires ou groupées en fascicules ou glomérules espacés le long du rachis.'Fleurs'sessiles ou pédicellées, à 1-3 bractées, parfois soudées entre elles; calice turbiniforme et soudé à l'ovaire dans sa partie inférieure, à 4-8 lobes généralement peu accrescents; pétales en même nombre que les sépales et alternant avec eux, imbriqués dans le bouton, parfois longuement accrescents et persistants; étamines opposées aux pétales, en même nombre que ceux-ci ou par fascicules de 3 (rarement plus), alternant avec des glandes subglobuleuses-aplaties ; filets filiformes ; anthères subglobuleuses, dorsifixes, extrorses, à 2 loges à déhiscence longitudinale; ovaire semi-infère, uniloculaire, à 2-6 placentas pariétaux pluriovulés; stigmates 2-6, ± soudés à la base et libres au sommet.'Capsules'déhiscentes en 2-6 valves, entourées du périanthe ± accrescent et persistant.'Graines peu nombreuses, petites, albuminées.\n\t\t\t\tGenre pantropical comprenant environ 140 espèces. En Afrique tropicale : 59 espèces réparties en 8 sections. Pour la Flore : 6 espèces appartenant à 2 sections.

    • Provided by: [C].Flore d'Afrique Centrale
      • Source: [
      • 14
      • ]. 

      Distribution

      A large pantropical genus of over 200 species

    • Provided by: [I].Flora de Panama
    • Included Species

       Information From

      MBG Floras Images
      http://www.tropicos.org/ImageSearch.aspx
      Flora images. Missouri Botanical Garden. Accessed on Jun. 2018.
      • A Missouri Botanical Garden
      • B Missouri Botanical Garden
      Flore d'Afrique Centrale
      https://www.floredafriquecentrale.be
      • C http://creativecommons.org/licenses/by/4.0/ Attribution 4.0 International (CC BY 4.0)
      Plants Of the World Online Portal - FTEA
      https://www.kew.org/science/who-we-are-and-what-we-do/strategic-outputs-2020/plants-of-the-world-online
      http://www.plantsoftheworldonline.org/terms-and-conditions
      • D
      Flora de Nicaragua
      http://www.tropicos.org/projectwebportal.aspx?projectid=7&pagename=Home&langid=66
      WD Stevens, CU Ulloa, A Pool and OM Montiel. Missouri Botanical Garden Press, 2001
      • E Missouri Botanical Garden
      Manual de Plantas de Costa Rica
      http://www.tropicos.org/Project/Costa%20Rica
      Hammel, B. E.; Grayum, M. H.; Herrera, C.; Zamora, N. Manual de Plantas de Costa Rica. Missouri Botanical Garden Press, 2003-2014
      • F Missouri Botanical Garden
      Flora Of CHina @ efloras.org
      http://www.efloras.org/flora_page.aspx?flora_id=2
      'Flora of China @ eFloras (2008). Published on the Internet http://www.efloras.org/flora_page.aspx?flora_id=2 [accessed August 2016]' Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
      • G Missouri Botanical Garden
      Plants Of the World Online Portal - FZ
      https://www.kew.org/science/who-we-are-and-what-we-do/strategic-outputs-2020/plants-of-the-world-online
      http://www.plantsoftheworldonline.org/terms-and-conditions
      • H
      Flora de Panama
      http://www.tropicos.org/Project/PAC
      Robert E. Woodson, Jr. and Robert W. Schery Annals of the Missouri Botanical Garden Vol. 67, No. 4 (1980), pp. ii-xxxiii
      • I Missouri Botanical Garden
      Salicaceae
      • J CC0 1.0 Universal (CC0 1.0).
      World Flora Online consortium
      http://www.worldfloraonline.org/organisation/WFO
      World Flora Online Data. 2018.
      • K CC0 1.0 Universal (CC0 1.0).