Humulus L.
  • Sp. Pl. 2: 1028. 1753. (1 May 1753)
  • Hop, houblon [Latin humulus, applied to hop plant]

Cite taxon page as 'WFO (2021): Humulus L. Published on the Internet; Accessed on: 27 Oct 2021'

General Information

Herbs, perennial or annual, twining, dioecious. Stems, branchlets, and petioles with rigid 2-armed stalked hairs. Stems coarse, 6-ridged or winged. Leaves opposite, well developed leaves ± cordate and 3-7(-9)-lobed, terminal leaves sometimes ± ovate and simple, abaxially with yellowish brown resinous glands and dots. Male inflorescences laxly paniculate. Male flowers: filaments straight in bud; female inflorescences a conelike spicate cyme; bracts imbricate, persistent and enlarged in fruit, margin entire. Female flowers: calyx thinly membranous, appressed to ovary, margin entire; ovary ± surrounded by appressed calyx; styles 2-branched; branches caducous. Achene broadly ovoid; calyx persistent, appressed to achene; pericarp crustaceous; embryo spirally involute; cotyledons narrow.

  • Provided by: [C].Flora Of CHina @
    • Source: [
    • 1
    • ]. 

    Herbs , taprooted annuals or rhizomatous perennials, rightward-twining. Stems usually branched, armed with rigid 2-branched, stalked hairs that facilitate climbing. Leaves simple; petioles often twining, with 2-branched hairs. Leaf blade mostly cordate, palmately lobed, sometimes unlobed; surfaces abaxially resin-dotted and×or gland-dotted. Inflorescences: staminate inflorescences axillary and terminal, cymose panicles, erect to pendent, (10-)20-100+-flowered, flowers small; pistillate axillary, spikes or racemes, flowers solitary or paired, short-pedicellate, subtended by bracts and bracteoles. Staminate and pistillate flowers usually on different plants. Achenes lenticular or terete, ensheathed by brownish or sometimes mottled persistent perianth; embryo coiled. x = 10.

  • Provided by: [B].Flora of North America @
    • Source: [
    • 2
    • ]. 

    "Dioecious; pistillate fls in short spikes, paired, each pair subtended by a foliaceous bract; ovary enclosed by the membranous, lobeless cal; fr an achene enclosed in the persistent cal and covered by the accrescent bracts; twining, herbaceous, perennial vines with rough stems and broad, opposite, usually lobed lvs. 3, N. Temp."

  • Provided by: [D].New York Botanical Garden
    • Source: [
    • 3
    • ]. 



    Small, E. 1978. A numerical and nomenclatural analysis of morpho-geographic taxa of Humulus. Syst. Bot. 3: 37-76. Small, E. 1980. The relationships of hop cultivars and wild variants of Humulus lupulus. Canad. J. Bot. 58: 676-686.

  • Provided by: [B].Flora of North America @
    • Source: [
    • 2
    • ]. 

    Other Local Names

    Hop, houblon [Latin humulus, applied to hop plant]

     Information From

    World Flora Online Data. 2017.
    • A CC0 1.0 Universal (CC0 1.0).
    Flora of North America @
    'Flora of North America @ eFloras (2008). Published on the Internet [accessed August 2016]' Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
    • B Flora of North America Association
    Flora Of CHina @
    'Flora of China @ eFloras (2008). Published on the Internet [accessed August 2016]' Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
    • C Missouri Botanical Garden
    New York Botanical Garden
    Descriptions of plants should be attributed to the full citation for each individual article, chapter or book that is the source for each record, which should include the authors of original publication.
    • D Content licensed under Creative Commons Attribution-NonCommercial-ShareAlike 3.0 Unported License
    World Flora Online consortium
    World Flora Online Data. 2018.
    • E CC0 1.0 Universal (CC0 1.0).