Herbs perennial, subshrubs, or shrubs, sometimes epiphytic or epilithic, without stinging hairs. Stems often succulent. Leaves apparently alternate, distichous; nanophylls often present, very small, opposite to developed leaves; stipules intrapetiolar, small, undivided; leaf blade ± fleshy, pinnately veined, base asymmetric, margin usually entire or serrulate; cystoliths linear. Inflorescences axillary, sometimes at leafless nodes, of unisexual flowers (plants monoecious or dioecious); male glomerules in lax cymes or crowded on a small, bractless receptacle; female capitula solitary, subsessile, on fleshy subglobose receptacle; bracteoles present, often spatulate. Male flowers: perianth lobes 4 or 5, segments often valvate; stamens 4 or 5; filaments inflexed in bud; rudimentary ovary globose or obovoid. Female flowers: perianth lobes 3 or 4, very small, boat-shaped, free or connate at base, ± fleshy; staminodes absent. Ovary straight; stigma sessile, penicillate, soon disappearing; ovule orthotropous. Achene ovoid or ellipsoid, not ribbed, enclosed by persistent, fleshy perianth or exserted from small perianth. Seeds without endosperm; cotyledons ovate.
Perennial herbs, sometimes shrubs, sometimes epiphytic, stems often rather juicy or succulent, monoecious or dioecious by abortion
Male inflorescences small pedunculate clusters; ♀ inflorescences capitate, with a fleshy subglobose receptacle on short fleshy peduncles, in the axils of fallen leaves
Leaves basically opposite, but one leaf of each pair is either reduced or early falling, thus the plants appear to have alternate leaves; lamina very unequal-sided at base, entire, crenate or serrate, rather fleshy; stipules intrapetiolar, fused to the apex
Male flowers 4–5-merous; rudimentary ovary present, obovoid
Achene ovate, partly covered by the fleshy persistent perianth.
Female flowers 4–5-merous, with very short perianth; ovary straight, exceeding the perianth; stigma sessile, penicillate
Perennial herbs, often epiphytic or epilithic, monoecious or dioecious by abortion; stems juicy or succulent.
Leaves distichous, opposite and heteromorphous with one leaf of each pair reduced in size, or leaves apparently alternate with one in each pair completely reduced; lamina crenate or serrate, rather fleshy, penninerved, with linear cystoliths.
Inflorescences unisexual, in the axils of existing or fallen leaves; male flowers in small pedunculate glomerules; female inflorescences capitate with fleshy receptacles on short fleshy peduncles.
Achene ovoid, surrounded at the base by the accrescent perianth.
Female flowers sessile with a persisting perianth of 4–5 free tepals; ovary erect, symmetrical, with a penicillate stigma.
Male flowers pedicellate, pentamerous (rarely tetramerous), with a rudimentary pistillode.
Herbes'charnues ou sous-arbustes, monoïques ou dioïques, à feuilles alternes (comme celles de'Elatostema ), distiques, asymétriques à la base, aiguës au sommet, penninerves, brièvement pétiolées; stipules intrapétiolaires, entières.'Inflorescences'sans involucre, solitaires ou fasciculées, axillaires sur la partie effeuillée de la tige; les ♂ en glomérules; les ♀ en capitules sur un réceptacle charnu.'Fleurs'♂ 4-5-mères; périgone membraneux, sans appendices; fleurs ♀ sessiles, à périgone 3-4-partite, très petit et persistant; ovaire dressé, à stigmate en pinceau.'Akènes ovoïdes.\n\t\t\t\tEnviron 10 espèces tropicales, d'Asie, de Polynésie et d'Afrique; 1 au Congo belge.