Seeds solitary in each locule by abortion, ellipsoid-ovoid, ecarunculate; albumen fleshy; cotyledons broad and flat.
Fruit globose, tardily septicidally dehiscent, subdrupaceous; exocarp smooth, shiny, becoming wrinkled on drying; mesocarp spongy; endocarp thick, bony or woody, 3-locular
Female flowers: sepals ± as in the ♂ flowers; disc cupular; ovary 3-locular, with 2 ovules per locule; styles 3, connate at the base, bifid
Male flowers: buds ovoid; sepals 5(–6), imbricate; petals 0; disc extrastaminal, annular, ± lobed, the lobes alternating with the sepals; stamens (4–)5–7, filaments connate, anthers introrse, thecae parallel, longitudinally dehiscent; pistillode columnar, often trifid
Flowers in axillary densely few-flowered pedunculate or subsessile bracteate dichasia, or else solitary, arising with or below/after the leaves
Leaves alternate, petiolate, stipulate, simple, entire, somewhat coriaceous, penninerved
Dioecious or rarely monoecious trees or shrubs with a simple indumentum
Leaves alternate, petiolate, stipulate; blades simple, entire, penninerved.
Flowers in axillary pedunculate or subsessile bracteate dense few-(up to 12-)flowered cymes, or females solitary, arising among or below the leaves.
Dioecious or rarely monoecious trees or shrubs.
Male flowers: buds ovoid; sepals 5(6), imbricate; petals absent; disk extrastaminal, annular, shallowly lobed, the lobes alternating with the sepals; stamens (4)5–7, united for half to two-thirds of their length into a column, anthers basifixed, introrse, thecae parallel, longitudinally dehiscent; pistillode (non-functional ovary) usually trifid.
Female flowers: pedicels bibracteolate; buds and sepals as in the male; disk cupular; ovary 3-locular, with 2 ovules per loculus; styles 3, connate at the base, bifid, the stigmas coiled backwards.
Fruit globose, subdrupaceous, tardily septicidally dehiscent after the fleshy layer has been eaten by frugivores, and when the sun has dried out the endocarp; exocarp smooth, becoming wrinkled and shiny on drying; mesocarp spongy; endocarp thickly bony or woody, trilocular.