Trees usually large, prominently buttressed. Bark usually fissured, flaky. Stipules caducous, large or small; leaf blade ± leathery, tertiary veins parallel, margin entire. Flowers in axillary or terminal lax cymose panicles; bracts persistent, caducous, or absent. Sepals with 3 outer larger than 2 inner. Petals white, yellow, or pink, usually pubescent. Stamens (12-)15 or 20-100; anthers ovoid, oblong, or panduriform; connective subulate-cuspidate or stout, club-shaped; valves equal or outer one slightly larger. Ovary ovoid, pubescent; style subulate; stigma entire or 3-toothed. Fruit usually 1-seeded, closely surrounded by thickened bases of accrescent calyx segments; sepals developed into lorate wings, outer 3 much larger than 2 inner.