Boraginaceae Juss.
  • Gen. Pl. 128. 1789. (4 Aug 1789)


Cite taxon page as 'WFO (2020): Boraginaceae Juss. Published on the Internet;http://www.worldfloraonline.org/taxon/wfo-7000000079. Accessed on: 10 Jul 2020'

General Information

Herbs perennial, biennial, or annual, less often lianas, shrubs, or trees, usually bristly or scabrous-pubescent. Leaves simple, exstipulate, alternate, rarely opposite, entire or serrate at margin. Inflorescences often double scorpioid cymes, rarely solitary; bracts present or absent. Flowers bisexual, actinomorphic, rarely zygomorphic. Calyx usually 5-parted or lobed, mostly persistent. Corolla tubular, campanulate, rotate, funnelform, or salverform; tube appendages 5, rarely more, mostly trapeziform, rarely absent, sometimes a ring of hairs present; limb usually 5-parted; lobes overlapping, rarely twisted in bud. Stamens 5, inserted on corolla tube or rarely at throat, included or rarely exserted; anthers introrse, 2-loculed, usually dorsifixed at base, less often medifixed, dehiscence longitudinal. Nectaries at base of corolla tube or on disc below ovary. Ovary superior, 2-carpellate; locules 2 and each with 2 ovules, or 4 and each with 1 ovule; ovules nearly atropous, semianatropous, or anatropous. Style terminal or gynobasic, branched or not. Gynobase flat, fastigiate, or subulate. Fruit 1-4-seeded drupes or nutlets (mericarps); nutlets mostly dry, often ornamented with wings, prickles and/or glochids (stiff bristles with barbed or anchorlike tips). Seeds vertical or oblique, coat membranous; embryo straight, less often curved; cotyledons flat, fleshy.

  • Provided by: [D].Flora Of CHina @ efloras.org
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    Morphology

    Fruit a drupe or of 4: nutlets

  • Provided by: [B].Plants Of the World Online Portal - FWTA
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    Seeds with or without endosperm

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    Ovary superior, 2-celled or 4-celled by spurious septa, entire or deeply 4-lobed; style terminal or gynobasic

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    Ovules paired, erect or spreading from the central axis

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    Anthers 2-celled, opening lengthwise

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    Disk present or absent

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    Corolla with contorted or imbricate lobes

  • Provided by: [B].Plants Of the World Online Portal - FWTA
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    • 1
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    Stamens the same number as the corolla-lobes and alternate with them, inserted on the corolla

  • Provided by: [B].Plants Of the World Online Portal - FWTA
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    Flowers often in scorpoid cymes, actinomorphic or rarely oblique, mostly hermaphrodite

  • Provided by: [B].Plants Of the World Online Portal - FWTA
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    Calyx-lobes imbricate or rarely valvate

  • Provided by: [B].Plants Of the World Online Portal - FWTA
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    • 1
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    Leaves alternate or rarely opposite, simple; stipules absent

  • Provided by: [B].Plants Of the World Online Portal - FWTA
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    Trees, shrubs or herbs, often roughly scabrid or hispid

  • Provided by: [B].Plants Of the World Online Portal - FWTA
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    Nectariferous disc usually present

  • Provided by: [C].Plants Of the World Online Portal - FTEA
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    Style usually 1, 2 in one genus or sometimes 4 in one species of >i>Cordia, terminal or gynobasic, entire or 2–4-fid or twice 2-fid; stigmas clavate, capitate or peltate or linear to narrowly foliaceous

  • Provided by: [C].Plants Of the World Online Portal - FTEA
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    Fruit drupaceous with 1, 4-celled endocarp or separating in 2 bilocular or 4 unilocular 1-seeded pyrenes or with 4 quite distinct nutlets

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    Ovary superior, entire or deeply 4-lobed, 2-locular with 2 ovules in each or with 4, 1-ovuled locules due to the development of false partitions

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    Ovules axile, anatropous, erect, horizontal, basal or rarely pendulous

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    Seed with or without endosperm, erect, oblique or horizontal; testa membranous; embryo straight or curved; cotyledons flat or plicate

  • Provided by: [C].Plants Of the World Online Portal - FTEA
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    Annual or perennial herbs, often woody at the base, subshrubs, trees or woody climbers, usually characteristically scabrid or hispid

  • Provided by: [C].Plants Of the World Online Portal - FTEA
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    Leaves simple, alternate or less often opposite, petiolate or sessile, entire or variously toothed or crenate; stipules absent

  • Provided by: [C].Plants Of the World Online Portal - FTEA
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    Inflorescences terminal or axillary, bifurcate or in raceme-like or panicled groups, basically dichotomous cymes, the branches frequently scorpioid and spirally inrolled when young; bracts present or absent

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    Flowers hermaphrodite or unisexual (Cordia), sessile or pedicellate, mostly 5-merous, regular or somewhat irregular, occasionally heterostylous

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    Calyx tubular or campanulate, persistent and sometimes accrescent in fruit, with (2–)4–7 (or more) imbricate or rarely valvate or open lobes

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    • 2
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    Stamens the same number and alternating with the corolla-lobes, exserted or included, epipetalous; anthers 2- thecous

  • Provided by: [C].Plants Of the World Online Portal - FTEA
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    Corolla tubular, campanulate, funnel-shaped or rotate with tube very reduced; lobes 3–16, imbricate or contorted (>i>Myosotis); throat often with scales, thickenings, folds or crests

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    Corolla (4)5(6)-lobed, imbricate or contorted in bud, tubular, funnel-shaped, campanulate, salver-shaped or rotate; tube sometimes with folds, scales or hairs in the throat

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    Stamens (or staminodes) as many as corolla lobes and alternate with them, inserted on the corolla tube; anthers 2-thecous, dehiscing longitudinally, introrse, dorsifixed

  • Provided by: [A].Plants Of the World Online Portal - FZ
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    Disk present or absent

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    Leaves alternate, rarely subopposite or opposite, simple exstipulate

  • Provided by: [A].Plants Of the World Online Portal - FZ
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    Inflorescence usually cymose, composed of one or more helicoid or scorpioid cymes, sometimes paniculate or racemose or sometimes flowers solitary, terminal or axillary

  • Provided by: [A].Plants Of the World Online Portal - FZ
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    Flowers hermaphrodite or sometimes unisexual, actinomorphic or sometimes zygomorphic

  • Provided by: [A].Plants Of the World Online Portal - FZ
    • Source: [
    • 3
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    Calyx (4)5(6)-lobed or 3–5-toothed, imbricate or valvate

  • Provided by: [A].Plants Of the World Online Portal - FZ
    • Source: [
    • 3
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    Herbs, shrubs or rarely trees

  • Provided by: [A].Plants Of the World Online Portal - FZ
    • Source: [
    • 3
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    Fruit drupaceous with a 1–4-seeded stone or splitting into 4 pyrenes, or of one to four-seeded nutlets; seeds generally without endosperm

  • Provided by: [A].Plants Of the World Online Portal - FZ
    • Source: [
    • 3
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    Ovary superior, entire or deeply 4-lobed, bi-locular or spuriously 4-locular, placentation axial, ovules 4, erect or spreading; style 1, terminal or gynobasic, entire or cleft or twice cleft, or styles 2, terminal (Coldenia)

  • Provided by: [A].Plants Of the World Online Portal - FZ
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    Included Genus

     Information From

    Plants Of the World Online Portal - FZ
    https://www.kew.org/science/who-we-are-and-what-we-do/strategic-outputs-2020/plants-of-the-world-online
    http://www.plantsoftheworldonline.org/terms-and-conditions
    • A
    Plants Of the World Online Portal - FWTA
    https://www.kew.org/science/who-we-are-and-what-we-do/strategic-outputs-2020/plants-of-the-world-online
    http://www.plantsoftheworldonline.org/terms-and-conditions
    • B The Trustees of the Royal Botanic Gardens, Kew
    Plants Of the World Online Portal - FTEA
    https://www.kew.org/science/who-we-are-and-what-we-do/strategic-outputs-2020/plants-of-the-world-online
    http://www.plantsoftheworldonline.org/terms-and-conditions
    • C
    Flora Of CHina @ efloras.org
    http://www.efloras.org/flora_page.aspx?flora_id=2
    'Flora of China @ eFloras (2008). Published on the Internet http://www.efloras.org/flora_page.aspx?flora_id=2 [accessed August 2016]' Missouri Botanical Garden, St. Louis, MO & Harvard University Herbaria, Cambridge, MA.
    • D Missouri Botanical Garden
    Boraginaceae
    World Flora Online Data. 2017.
    • E CC0 1.0 Universal (CC0 1.0).
    World Flora Online consortium
    http://www.worldfloraonline.org/organisation/WFO
    World Flora Online Data. 2018.
    • F CC0 1.0 Universal (CC0 1.0).