Shrubs or trees, to 12 m. Leaves alternate (distichous), petiolate, stipitate; stipules subulate or filiform [absent or peltate discs]; blade palmately veined, seldom lobed, base often asymmetric, margins serrate, surfaces hairy, hairs usually mixed: unicellular and multicellular, simple with some setiform and some glandular, branched, and stellate, often ± tangled, forming tomentum. Inflorescences: flowers solitary or in clusters of 2–3+, usually supra-axillary; involucel absent [bracteoles ca. 15, filiform]. Flowers: sepals caducous [persistent], (4–)5(–7), valvate, basally distinct or weakly connate; petals caducous, (4–)5(–7), distinct; nectaries absent; stamens 10–75+, filaments distinct or bases connate; ovary superior [inferior], 5–7-carpellate; style 1 ; stigmas 5–7, ± decurrent. Fruits baccate, ± spheric. Seeds [25–]100–200+.
Stamens numerous with filiform filaments, free or nearly so; anthers of 2 thecae, sometimes versatile, fixed at or near the base of the connective, dehiscing by longitudinal slits, sometimes only apically
Ovary superior to inferior, syncarpous, 5–many locular but sometimes unilocular at apex; placentae (where known) lobed, pendulous with numerous anatropous ovules; style thick, or sometimes ± lacking; stigma thick, lobed-sulcate
Fruit a berry with numerous small seeds
Leaves alternate, distichous, petiolate, cordate at the base, palmately nerved, serrate; stipules (where known) filiform or leafy and peltate
Inflorescences supra-axillary with 1–few pedicellate usually hermaphrodite flowers in clusters
Sepals (4–)5(–7), valvate, with ± spreading tips, joined at base to form a saucer-shaped or cupular tube
Petals (4–)5(–7), free, imbricate and ± crumpled in bud, often exceeding the calyx, thin, falling