Annuals or perennials, affixed or suspended aquatic. Rhizoids absent or present, filiform, branched. Stolons filiform, much branched, often mat-forming. Traps lateral on leaf segments, stalked, ovoid, 1-2.5 mm, mouth lateral; appendages 2, dorsal, branched, setiform, with shorter setae. Leaves numerous on stolons, 0.5-1.5 cm; primary segments (1 or)2, unbranched or sparsely dichotomously branched into 3-8 ultimate segments; ultimate segments capillary, slightly flattened, margin entire or sparsely denticulate, apex and teeth setulose. Inflorescences erect, 2-15 cm, 1-3(-6)-flowered; peduncle terete, 0.3-0.5 mm thick, glabrous; scale 1, similar to bracts; bracts basifixed, semiorbicular, ca. 1 mm, minutely glandular, apex truncate and obscurely dentate. Pedicel erect to spreading, 2-12 mm, filiform; bracteoles absent. Calyx lobes subequal, broadly ovate to orbicular, 1.5-2 mm, apex rounded. Corolla yellow, 4-8 mm; lower lip slightly smaller than upper lip, base with a prominent 2-lobed swelling, apex rounded; spur narrowly conic to cylindric from a conic base, shorter or longer than corolla lower lip, distal part sparsely stipitate glandular, apex obtuse; palate densely pubescent; upper lip broadly ovate to suborbicular, ca. 2 × as long as upper calyx lobe, apex obscurely 3-lobed. Filaments 1-1.5 mm, curved; anther thecae confluent. Ovary globose; style evident; stigma lower lip transversely elliptic, upper lip obsolete. Capsule globose, 2-3 mm in diam., 2-valvate. Seeds lenticular, 0.8-1 mm in diam., margin broadly winged, wing shallowly and irregularly dentate; seed coat with small prominent reticulations. Fl. Apr-Nov, fr. May-Dec. 2n = 28. Utricularia gibba L., Sp. Pl. 18. 1753; U. obtusa Sw. Anuales o perennes, acuáticas. Hojas divididas, hasta 15 mm de largo, con trampas laterales en los segmentos. Racimos 120 cm de largo, pedicelos 230 mm de largo; lobos del cáliz subiguales, orbiculares, 13 mm de largo; corola 425 mm de largo, amarilla, el labio inferior transversalmente elíptico, más largo a considerablemente más corto que el espolón. Cápsula globosa, 23 mm de diámetro, lateralmente 2-valvada. Poco común en aguas poco profundas o en lodo en las zonas pacífica y atlántica; 01000 m; fl y fr durante todo el año; Haynes 8318, Stevens 21523; trópicos y subtrópicos del mundo. Acuática, sumergida o varada en el barro; estolones numerosos, 0.2 –1 mm de ancho, cilíndricos, muy ra- mificados, que a menudo forman esteras enredadas; tubérculos ausentes; trampas sobre las hojas, relativamente pocas, 1–1.5 mm. Hojas numerosas, alternas, sésiles o pecioladas, 0.5 –1.5 cm, sumamente disecadas, los seg- mentos primarios usualmente 2, dicotómicamente ramificados, con segmentos capilares. Infls. erectas, emer- gentes (o a veces con fls. cleistógamas sumergidas), 1– 20(– 30) cm, con brácteas basifijas ca. 0.1 cm, semiorbiculares. Fls. principalmente 1 ó 2; cáliz con los lóbulos subiguales; corola amarilla (a menudo con una raya rojiza en el paladar del labio inferior), (3 –)4 – 6 mm, glabra (excepto por el paladar densamente pubes- cente), el labio superior ampliamente ovado a suborbicular, indistintamente (a veces distintamente) trilobulado en el ápice, el labio inferior ligeramente más pequeño, entero; espolón recto a ligeramente curvado hacia arriba, ligeramente más corto a más largo que el labio inferior, ± angostamente cónico, obtuso en el ápice. Frs. 2–3 mm de diám., globosos. Creeping stems delicate, seldom over 10 cm, forming mats near the bottom in shallow water; lvs scattered, rarely over 5 mm, commonly once (twice) forked, with a few bladders; flowering branches 5–10 cm, with 1–3 yellow fls 5–6 mm; palate prominent; spur very thick and obtuse, only ca half as long as the lower lip. Shallow water; Que. to Wis., s. to Fla. and La.; also Pacific states, W.I., and c. Amer. Hoja: forma capilar dicotómica(s); forma de las lámina(s) no aplicable; nervura(s) uninervea(s); inserción(es) del pecíolo(s) no aplicable; forma del utrículo(s) ( trampa ) ovoide(s); posición de la abertura del utrículo(s) lateral(es); apéndice(s) y posición de los apéndice(s) en el utrículo(s) dorsal(es); tipo de apéndice(s) del utrículo(s) ramificada(s)/recto(s). Inflorescencia: escama(s) y bractea(s) basifija(s)/base no auriculada(s)/margen(es) entera; bracteola(s) ausente(s). Flor: forma de la sépala(s) superior(es) oboval(es); forma de la sépala(s) inferior(es) circular(es); margen(es) del cáliz entera; nervura(s) del cáliz inconspicua(s); color de la corola amarilla; palato de la corola giboso(s)/de color amarilla o naranja; labio(s) inferior(es) de la corola inteiro/crenado(s); cálcar cónico(s)/encorvado(s)/recto(s)/más corto que el labio(s) inferior(es)/de igual dimensiones del que el labio(s) inferior(es)/más largo que el labio(s) inferior(es)/ápice(s) obtuso(s)/ápice(s) emarginado(s). Fruto: deiscencia bivalvar. Semilla: forma lenticular(es). Folha: forma capilar dicotômica(s); forma das lâmina(s) não aplicável; nervura(s) uninérvea(s); inserção(ções) do pecíolo(s) não aplicável; forma do utrículo(s) ( armadilha ) ovoide(s); posição da abertura do utrículo(s) lateral(ais); apêndice(s) e posição dos apêndice(s) no utrículo(s) dorsal(ais); tipo de apêndice(s) do utrículo(s) ramificada(s)/reto(s). Inflorescência: escama(s) e bráctea(s) basifixa(s)/base não auriculada(s)/margem(ns) inteira; bractéola(s) ausente(s). Flor: forma da sépala(s) superior(es) oboval(ais); forma da sépala(s) inferior(es) circular(es); margem(ns) do cálice(s) inteira; nervura(s) do cálice(s) inconspícua(s); cor da corola amarela; palato da corola giboso(s)/de cor amarela ou laranja; lábio(s) inferior(es) da corola inteiro/crenado(s); cálcar cônico(s)/curvado(s)/reto(s)/mais curto que o lábio(s) inferior(es)/de igual tamanho do que o lábio(s) inferior(es)/mais longo que o lábio(s) inferior(es)/ápice(s) obtuso(s)/ápice(s) emarginado(s). Fruto: deiscência bivalvar. Semente: forma lenticular(es). Leaf: form capillary dichotomous; form of the blade non applicable; veins uninerveous; insertion of the petiole not applicable; form of the utricle(s) ( trap ) ovoid; position of the opening of the utricle(s) lateral; appendix and position of the appendix on the utricle(s) dorsal; type of the appendix of the utricle(s) branched/straight. Inflorescence: scale and bract(s) basifixed/base not auriculate/margin entire; bracteole(s) absent. Flower: form of the sepal upper obovate; form of the sepal lower circular; margin of the calyx entire; veins of the calyx inconspicuous; colour of the corolla yellow; palate of the corolla gibbose/of the colour yellow or orange; lip lower of the corolla entire/crenate; spur conical/bent/straight/shorter than the lip lower/of the equal size of the than the lip lower/longer than the lip lower/apex obtuse/apex emarginate. Fruit: dehiscence bivalve. Seed: form lenticular. Inflorescence erect, 2–35 cm. high, solitary or often several arising in succession from the fascicle of stolons; flowers (1)2(6); peduncle straight, smooth and glabrous; scales absent or more usually 1 at or above the middle of the peduncle, similar to the bracts; bracts basifixed, semi-circular, semi-amplexicaul, more or less 1 mm. long; bracteoles absent; pedicels filiform, erect, (2)6–12(30) mm. long. Filaments linear; anther-thecae more or less confluent. Ovary globose; style short but distinct; stigma inferior lip semi-circular, superior short, more or less obsolete. Calyx lobes subequal, circular to broadly ovate, 1–3 mm. long. Corolla yellow, often with brown or reddish nerves, 4–25 mm. long; superior lip circular, 2–3 times as long as the upper calyx lobe, usually shallowly and obscurely 3-lobed; inferior lip usually shorter and narrower than the superior, circular, usually entire, rarely emarginate; palate much raised and bigibbous; spur conical to narrowly cylindrical, shorter than to almost twice as long as the inferior lip, the apex usually bearing a few shortly stalked glands. Seeds rather few (more or less 20–40), imbricate on the smooth placenta, lenticular with a broad irregular wing, 1–1.6 mm. in total diam., distal surface smooth to verrucose; hilum prominent; testa-cells small, irregular, indistinct. Capsule globose, 2–4 mm. in diam., dehiscing into 2 lateral valves. Aquatic or subaquatic herb. Rhizoids few from the base of the peduncle, often absent, filiform. Stolons usually up to more or less 10, fasciculate at the base of the peduncle, filiform, terete, up to 20 cm.×1 mm., sometimes slightly inflated. Traps usually numerous, replacing one of the leaf segments at a fork, ovoid, 1–1.5 mm. long, stalked; mouth lateral, with 2 dorsal, long, usually copiously branched hairs and a few ventral shorter simple hairs. Leaves numerous, alternate on the stolons, up to 20 mm. long; segments capillary, glabrous or sparsely setulose, simple or usually dichotomously forked from or near the base, each segment sometimes again 1–3 or rarely more times forked. Inflorescence erect, 2–35 cm. high, solitary or often several arising in succession from the fascicle of stolons; flowers (1–)2(–6); scape straight, smooth and glabrous; scales absent or more usually 1 at or above the middle of the scape, similar to the bracts; bracts basifixed, semi-orbicular, semi-amplexicaul, ± 1 mm. long; bracteoles absent; pedicels filiform, erect, (2–)6–12(–30) mm. long. Traps usually numerous, replacing one of the leaf-segments at a fork, ovoid, 1–1.5 mm. long, stalked; mouth lateral; upper lip with 2 long usually copiously branched hairs; lower lip usually with a few shorter simple hairs. Corolla yellow, often with brown or reddish nerves, 4–25 mm. long; upper lip orbicular, 2–3 times as long as the upper calyx-lobe, usually shallowly and obscurely 3-lobed; lower lip usually shorter and narrower than the upper, orbicular, usually entire, rarely emarginate; palate much raised and bigibbous; spur conical to narrowly cylindrical, shorter than to almost twice as long as the lower lip, apex usually bearing a few shortly stalked glands. Calyx-lobes subequal, orbicular to broadly ovate, 1–3 mm. long. Aquatic or subaquatic herb. Stolons usually up to ± 10, fasciculate at the base of the scape, filiform, terete, up to 20 cm. long or more, sometimes slightly inflated, up to 1 mm. thick. Rhizoids few from the base of the scape, often absent, filiform. Leaves numerous, alternate on the stolons, up to 20 mm. long; segments capillary, glabrous, simple or usually dichotomously forked from or near the base, each segment sometimes again 1–3 or rarely more times forked. Ovary globose; style short but distinct; stigma lower lip semi-orbicular, upper short, ± obsolete. Capsule globose, 2–4 mm. in diameter, dehiscing into 2 lateral valves. Filaments linear; anther-thecae ± confluent. Seeds rather few (± 20–40), imbricate on the smooth placenta, lenticular with a broad irregular wing, 1–1.6 mm. in total diameter, outer surface smooth to verrucose; hilum prominent; testa-cells small, irregular, indistinct. Small to medium sized, annual or perennial, affixed or suspended aquatic. Rhizoids absent or present, filiform, a few cm long, 0.2-0.4 mm thick, bearing a few very short, dichotomously divided branches with curved, papillose, apically obtuse segments; sometimes the rhizoids with an indefinite apical transformation as a stolon. Stolons usually numerous, much branched and often mat-forming, filiform, terete, glabrous, up to 20 cm long or longer, 0.2-1 mm thick, sometimes somewhat thicker and spongy. Leaves numerous, 0.5-1.5 cm long, ± uniform, the primary segments 2 (or sometimes reduced to 1), sparsely dichotomously branched into up to 8 (but usually not more than 4) ultimate segments, or sometimes 1 branch of each dichotomy replaced by a trap, the ultimate segments capillary, slightly flattened, laterally entire or sparsely denticulate, the apex of the segments and of the lateral teeth when present setulose. Traps rather few, lateral on the primary to penultimate segments, ovoid, stalked, 1-2.5 mm long, the mouth lateral with 2 dorsal, setiform, branched appendages and further dorsal, lateral and ventral setae, the internal glands 2- and 4-armed, the arms narrowly ovoid, with apex acute, 10-40 µ long, 1.5-6 times as long as wide, the quadrifids with both pairs ± equally slightly divergent. Inflorescence erect, emergent, (or sometimes submersed, much reduced and with cleistogamous flowers), 1-20(-30) cm long, solitary or frequently 2 or more arising together or in succession; peduncle filiform, terete, glabrous, ± straight, 0.4-0.8 mm thick. Scales usually 1, sometimes 2, at or near the middle of the peduncle, similar to the bract, or frequently absent. Bracts basifixed, semi-amplexicaul, approximately semicircular, with apex rounded, truncate or obscurely dentate, c. 1 mm long, minutely glandular, obscurely nerved. Bracteoles absent. Flowers (1-)2-6(-12), the raceme axis short to elongate; pedicels erect or spreading, filiform, terete, 0.2-3 cm long. Calyx lobes subequal, broadly ovate or circular with apex rounded, 1-3 mm long. Corolla yellow, often with reddish-brown nerves, 4-25 mm long; upper lip broadly ovate to almost circular, usually ± obscurely to distinctly 3-lobed; lower lip limb usually slightly smaller, circular to transversely elliptic, the base with a prominent, rounded, bilobed swelling, the apex rounded, entire; palate densely pubescent with simple unicellular hairs; spur narrowly conical or narrowly cylindrical from a conical base with apex obtuse or ± obscurely bifid, a little shorter than to distinctly longer than the lower lip, the distal part externally, sparsely, minutely stipitate-glandular. Filaments curved, 1-1.5 mm long, the anther thecae ± confluent. Ovary globose; style distinct; stigma lower lip transversely elliptic, the upper lip obsolete. Capsule globose, 2-3 mm in diameter, the wall membranous, tardily laterally bivalvate. Seeds imbricate on the placenta, thickly lenticular, approximately circular in dorsal view, 0.8-1 mm in diameter, with a relatively broad, thick wing, the wing produced to form a rounded lobe on that part of the margin which, in situ on the placenta, is nearest the apex of the capsule, the rest of the margin shallowly and irregularly dentate, the ventral surface smooth or sometimes with multicellular, obtusely conical processes, the testa cells small, slightly elongate, with the anticlinal boundaries sunken, the periclinal walls smooth, convex, but with the central area collapsing and becoming concave. Flowering more or less throughout the year when conditions are suitable. Submerged, aquatic annual or perennial to 20 cm. Leaves sparsely dissected. Flowers in an ebracteolate, emergent raceme, yellow, lips subequal, broadly ovate, palate obscuring mouth, spur ± as long as or longer than lower lip. Fruits erect. Aquatic or subaquatic herb. Leaves with a few segments, 5-15 mm long. Flowers: bracteoles 0, inflorescence 1-4-flowered, without whorl of inflated leaf-like organs; corolla with spur well developed, 4-25 mm long, not hairy outside; yellow, often with brown or reddish nerves; Oct.-Jun. Traps ovoid, 1.0-1.5 mm long. Terrícola Epífita Aquática Shallow still or slowly flowing water, or in mud in ditches, pools, lakes, river backwaters, bogs, swamps and marshes. Also frequently in deep water but not flowering unless supported on floating mats of living or dead vegetation. Usually at low altitudes but ascending to 2500 m alt. Boggy or shallow water. Dry Grassland. Shallow water and mud by ditches, streams and lakes. Pan tropical, extending northwards into eastern and western Canada and U.S.A., Spain and Portugal, North Africa, Israel, China and Japan and southwards to Argentina, S. Africa, Australia and New Zealand, also as an occasional introduction into other European countries and a common weed of aquaria and botanic gardens throughout the world. America: Canada (Nova Scotia, New Brunswick, Quebec, Ontario, [Manitoba], British Colombia), U.S.A.! (absent only in the great plains and Rocky Mountains states i.e. N & S Dakota, Nebraska, Iowa, Montana, Wyoming, Colorado, New Mexico, Arizona, Utah, Idaho, Nevada, and also Alaska, but in Hawaii), Mexico (Coahuila, Durango, Tamaulipas, San Luis Potosi, Jalisco, Tabasco, Michoacan, Mexico, Puebla, Chiapas, Campeche, Vera Cruz, Yucatan, and very probably elsewhere), Guatemala, El Salvador, Honduras, Belize, Nicaragua, Costa Rica, Panama, Bahamas, Cuba, Jamaica, Haiti, Dominican Rep., Puerto Rico, Antigua, Guadaloupe, Marie Galant, Dominca, Martinique, Trinidad!, Venezuela (Sucre, Monagas, Delta Amacuro, Anzoategui, Guarico, Miranda, Bolivar, Amazonas, Apure, Barinas, Portugesa, Cojedes, Carabobo, Falcon, Aragua, Yaracuy, Merida, Zulia), Colombia (Santander, Cundinamarca, Cauca, Amazonas), Guyana, Surinam, French Guiana, Brazil (Para, Amazonas, Maranhão, Ceara, Piaui, Pernambuco, Bahia, Goias, Mato Grosso, Minas Gerais, Espirito Santo, Guanabara, Rio de Janeiro, Sao Paulo, Sta. Catarina, Parana, Rio Grande do Sul), Ecuador, Peru, Bolivia, Paraguay, Uraguay, Argentina (Chaco, Jujuy, Salta, Tucuman, Corrientes, Misiones, Santa Fe, Cordoba, Buenos Aires). Europe: England (introduced), Portugal, Spain, Hungary (introduced). N. Africa: Morocco, Algeria, Egypt. Tropical and S. Africa: Senegal, Mali, Guinea-Bissau, Guinée, Sierra Leone, Liberia, Côte d’Ivoire, Burkina, Ghana, Nigeria, Niger, Cameroun, Central African Republic, Chad, Gabon, Congo, Zaïre, Sudan, Ethiopia, Uganda, Kenya, Rwanda, Tanzania!, Zambia, Zimbabwe, Moçambique, Angola, Namibia, Botswana, Transvaal, Natal, Cape Province, Madagascar, Mauritius. Asia: Israel, India (Rajasthan, Uttar Pradesh, Maharashtra, Madhya Pradesh, Andhra Pradesh, Karnataka, Tamil Nadu, Kerala, W. Bengal, Meghalaya, [Nicobar Islands]), Nepal, Ceylon, Burma, Thailand, Vietnam, Peninsular Malaysia, Sumatra, Java, Borneo, Philippines, Sulawesi, New Guinea, China (Yunnan, Guanxi, Guangdong, Hong Kong, Hunan, Henan, Fujian, Taiwan), Japan (southern), Palau. Australasia: Australia (Western Australia!, New South Wales, Queensland!, Northern Territory!), New Zealand (possibly introduced), New Caledonia. Uitenhage and throughout the tropics. Acuática, sumergida o varada en el barro; estolones numerosos, 0.2 –1 mm de ancho, cilíndricos, muy ra- mificados, que a menudo forman esteras enredadas; tubérculos ausentes; trampas sobre las hojas, relativamente pocas, 1–1.5 mm. Hojas numerosas, alternas, sésiles o pecioladas, 0.5 –1.5 cm, sumamente disecadas, los seg- mentos primarios usualmente 2, dicotómicamente ramificados, con segmentos capilares. Infls. erectas, emer- gentes (o a veces con fls. cleistógamas sumergidas), 1– 20(– 30) cm, con brácteas basifijas ca. 0.1 cm, semiorbiculares. Fls. principalmente 1 ó 2; cáliz con los lóbulos subiguales; corola amarilla (a menudo con una raya rojiza en el paladar del labio inferior), (3 –)4 – 6 mm, glabra (excepto por el paladar densamente pubes- cente), el labio superior ampliamente ovado a suborbicular, indistintamente (a veces distintamente) trilobulado en el ápice, el labio inferior ligeramente más pequeño, entero; espolón recto a ligeramente curvado hacia arriba, ligeramente más corto a más largo que el labio inferior, ± angostamente cónico, obtuso en el ápice. Frs. 2–3 mm de diám., globosos. Annuals or perennials, affixed or suspended aquatic. Rhizoids absent or present, filiform, branched. Stolons filiform, much branched, often mat-forming. Traps lateral on leaf segments, stalked, ovoid, 1-2.5 mm, mouth lateral; appendages 2, dorsal, branched, setiform, with shorter setae. Leaves numerous on stolons, 0.5-1.5 cm; primary segments (1 or)2, unbranched or sparsely dichotomously branched into 3-8 ultimate segments; ultimate segments capillary, slightly flattened, margin entire or sparsely denticulate, apex and teeth setulose. Inflorescences erect, 2-15 cm, 1-3(-6)-flowered; peduncle terete, 0.3-0.5 mm thick, glabrous; scale 1, similar to bracts; bracts basifixed, semiorbicular, ca. 1 mm, minutely glandular, apex truncate and obscurely dentate. Pedicel erect to spreading, 2-12 mm, filiform; bracteoles absent. Calyx lobes subequal, broadly ovate to orbicular, 1.5-2 mm, apex rounded. Corolla yellow, 4-8 mm; lower lip slightly smaller than upper lip, base with a prominent 2-lobed swelling, apex rounded; spur narrowly conic to cylindric from a conic base, shorter or longer than corolla lower lip, distal part sparsely stipitate glandular, apex obtuse; palate densely pubescent; upper lip broadly ovate to suborbicular, ca. 2 × as long as upper calyx lobe, apex obscurely 3-lobed. Filaments 1-1.5 mm, curved; anther thecae confluent. Ovary globose; style evident; stigma lower lip transversely elliptic, upper lip obsolete. Capsule globose, 2-3 mm in diam., 2-valvate. Seeds lenticular, 0.8-1 mm in diam., margin broadly winged, wing shallowly and irregularly dentate; seed coat with small prominent reticulations. Fl. Apr-Nov, fr. May-Dec. 2n = 28. Hoja: forma capilar dicotómica(s); forma de las lámina(s) no aplicable; nervura(s) uninervea(s); inserción(es) del pecíolo(s) no aplicable; forma del utrículo(s) ( trampa ) ovoide(s); posición de la abertura del utrículo(s) lateral(es); apéndice(s) y posición de los apéndice(s) en el utrículo(s) dorsal(es); tipo de apéndice(s) del utrículo(s) ramificada(s)/recto(s). Inflorescencia: escama(s) y bractea(s) basifija(s)/base no auriculada(s)/margen(es) entera; bracteola(s) ausente(s). Flor: forma de la sépala(s) superior(es) oboval(es); forma de la sépala(s) inferior(es) circular(es); margen(es) del cáliz entera; nervura(s) del cáliz inconspicua(s); color de la corola amarilla; palato de la corola giboso(s)/de color amarilla o naranja; labio(s) inferior(es) de la corola inteiro/crenado(s); cálcar cónico(s)/encorvado(s)/recto(s)/más corto que el labio(s) inferior(es)/de igual dimensiones del que el labio(s) inferior(es)/más largo que el labio(s) inferior(es)/ápice(s) obtuso(s)/ápice(s) emarginado(s). Fruto: deiscencia bivalvar. Semilla: forma lenticular(es). Folha: forma capilar dicotômica(s); forma das lâmina(s) não aplicável; nervura(s) uninérvea(s); inserção(ções) do pecíolo(s) não aplicável; forma do utrículo(s) ( armadilha ) ovoide(s); posição da abertura do utrículo(s) lateral(ais); apêndice(s) e posição dos apêndice(s) no utrículo(s) dorsal(ais); tipo de apêndice(s) do utrículo(s) ramificada(s)/reto(s). Inflorescência: escama(s) e bráctea(s) basifixa(s)/base não auriculada(s)/margem(ns) inteira; bractéola(s) ausente(s). Flor: forma da sépala(s) superior(es) oboval(ais); forma da sépala(s) inferior(es) circular(es); margem(ns) do cálice(s) inteira; nervura(s) do cálice(s) inconspícua(s); cor da corola amarela; palato da corola giboso(s)/de cor amarela ou laranja; lábio(s) inferior(es) da corola inteiro/crenado(s); cálcar cônico(s)/curvado(s)/reto(s)/mais curto que o lábio(s) inferior(es)/de igual tamanho do que o lábio(s) inferior(es)/mais longo que o lábio(s) inferior(es)/ápice(s) obtuso(s)/ápice(s) emarginado(s). Fruto: deiscência bivalvar. Semente: forma lenticular(es). Leaf: form capillary dichotomous; form of the blade non applicable; veins uninerveous; insertion of the petiole not applicable; form of the utricle(s) ( trap ) ovoid; position of the opening of the utricle(s) lateral; appendix and position of the appendix on the utricle(s) dorsal; type of the appendix of the utricle(s) branched/straight. Inflorescence: scale and bract(s) basifixed/base not auriculate/margin entire; bracteole(s) absent. Flower: form of the sepal upper obovate; form of the sepal lower circular; margin of the calyx entire; veins of the calyx inconspicuous; colour of the corolla yellow; palate of the corolla gibbose/of the colour yellow or orange; lip lower of the corolla entire/crenate; spur conical/bent/straight/shorter than the lip lower/of the equal size of the than the lip lower/longer than the lip lower/apex obtuse/apex emarginate. Fruit: dehiscence bivalve. Seed: form lenticular. Folha: forma capilar dicotômica(s); forma das lâmina(s) não aplicável; nervura(s) uninérvea(s); inserção(ções) do pecíolo(s) não aplicável; forma do utrículo(s) ( armadilha ) ovoide(s); posição da abertura do utrículo(s) lateral(ais); apêndice(s) e posição dos apêndice(s) no utrículo(s) dorsal(ais); tipo de apêndice(s) do utrículo(s) ramificada(s)/reto(s). Inflorescência: escama(s) e bráctea(s) basifixa(s)/base não auriculada(s)/margem(ns) inteira; bractéola(s) ausente(s). Flor: forma da sépala(s) superior(es) oboval(ais); forma da sépala(s) inferior(es) circular(es); margem(ns) do cálice(s) inteira; nervura(s) do cálice(s) inconspícua(s); cor da corola amarela; palato da corola giboso(s)/de cor amarela ou laranja; lábio(s) inferior(es) da corola inteiro/crenado(s); cálcar cônico(s)/curvado(s)/reto(s)/mais curto que o lábio(s) inferior(es)/de igual tamanho do que o lábio(s) inferior(es)/mais longo que o lábio(s) inferior(es)/ápice(s) obtuso(s)/ápice(s) emarginado(s). Fruto: deiscência bivalvar. Semente: forma lenticular(es). Leaf: form capillary dichotomous; form of the blade non applicable; veins uninerveous; insertion of the petiole not applicable; form of the utricle(s) ( trap ) ovoid; position of the opening of the utricle(s) lateral; appendix and position of the appendix on the utricle(s) dorsal; type of the appendix of the utricle(s) branched/straight. Inflorescence: scale and bract(s) basifixed/base not auriculate/margin entire; bracteole(s) absent. Flower: form of the sepal upper obovate; form of the sepal lower circular; margin of the calyx entire; veins of the calyx inconspicuous; colour of the corolla yellow; palate of the corolla gibbose/of the colour yellow or orange; lip lower of the corolla entire/crenate; spur conical/bent/straight/shorter than the lip lower/of the equal size of the than the lip lower/longer than the lip lower/apex obtuse/apex emarginate. Fruit: dehiscence bivalve. Seed: form lenticular. Terrícola Epífita Aquática Epífita Aquática Erva Utricularia gibba L., Sp. Pl. 18. 1753; U. obtusa Sw. Anuales o perennes, acuáticas. Hojas divididas, hasta 15 mm de largo, con trampas laterales en los segmentos. Racimos 120 cm de largo, pedicelos 230 mm de largo; lobos del cáliz subiguales, orbiculares, 13 mm de largo; corola 425 mm de largo, amarilla, el labio inferior transversalmente elíptico, más largo a considerablemente más corto que el espolón. Cápsula globosa, 23 mm de diámetro, lateralmente 2-valvada. Poco común en aguas poco profundas o en lodo en las zonas pacífica y atlántica; 01000 m; fl y fr durante todo el año; Haynes 8318, Stevens 21523; trópicos y subtrópicos del mundo. Creeping stems delicate, seldom over 10 cm, forming mats near the bottom in shallow water; lvs scattered, rarely over 5 mm, commonly once (twice) forked, with a few bladders; flowering branches 5–10 cm, with 1–3 yellow fls 5–6 mm; palate prominent; spur very thick and obtuse, only ca half as long as the lower lip. Shallow water; Que. to Wis., s. to Fla. and La.; also Pacific states, W.I., and c. Amer. Inflorescence erect, 2–35 cm. high, solitary or often several arising in succession from the fascicle of stolons; flowers (1–)2(–6); scape straight, smooth and glabrous; scales absent or more usually 1 at or above the middle of the scape, similar to the bracts; bracts basifixed, semi-orbicular, semi-amplexicaul, ± 1 mm. long; bracteoles absent; pedicels filiform, erect, (2–)6–12(–30) mm. long. Traps usually numerous, replacing one of the leaf-segments at a fork, ovoid, 1–1.5 mm. long, stalked; mouth lateral; upper lip with 2 long usually copiously branched hairs; lower lip usually with a few shorter simple hairs. Corolla yellow, often with brown or reddish nerves, 4–25 mm. long; upper lip orbicular, 2–3 times as long as the upper calyx-lobe, usually shallowly and obscurely 3-lobed; lower lip usually shorter and narrower than the upper, orbicular, usually entire, rarely emarginate; palate much raised and bigibbous; spur conical to narrowly cylindrical, shorter than to almost twice as long as the lower lip, apex usually bearing a few shortly stalked glands. Calyx-lobes subequal, orbicular to broadly ovate, 1–3 mm. long. Aquatic or subaquatic herb. Stolons usually up to ± 10, fasciculate at the base of the scape, filiform, terete, up to 20 cm. long or more, sometimes slightly inflated, up to 1 mm. thick. Rhizoids few from the base of the scape, often absent, filiform. Leaves numerous, alternate on the stolons, up to 20 mm. long; segments capillary, glabrous, simple or usually dichotomously forked from or near the base, each segment sometimes again 1–3 or rarely more times forked. Ovary globose; style short but distinct; stigma lower lip semi-orbicular, upper short, ± obsolete. Capsule globose, 2–4 mm. in diameter, dehiscing into 2 lateral valves. Filaments linear; anther-thecae ± confluent. Seeds rather few (± 20–40), imbricate on the smooth placenta, lenticular with a broad irregular wing, 1–1.6 mm. in total diameter, outer surface smooth to verrucose; hilum prominent; testa-cells small, irregular, indistinct. Traps usually numerous, replacing one of the leaf-segments at a fork, ovoid, 1–1.5 mm. long, stalked; mouth lateral; upper lip with 2 long usually copiously branched hairs; lower lip usually with a few shorter simple hairs. Corolla yellow, often with brown or reddish nerves, 4–25 mm. long; upper lip orbicular, 2–3 times as long as the upper calyx-lobe, usually shallowly and obscurely 3-lobed; lower lip usually shorter and narrower than the upper, orbicular, usually entire, rarely emarginate; palate much raised and bigibbous; spur conical to narrowly cylindrical, shorter than to almost twice as long as the lower lip, apex usually bearing a few shortly stalked glands. Calyx-lobes subequal, orbicular to broadly ovate, 1–3 mm. long. Aquatic or subaquatic herb. Stolons usually up to ± 10, fasciculate at the base of the scape, filiform, terete, up to 20 cm. long or more, sometimes slightly inflated, up to 1 mm. thick. Rhizoids few from the base of the scape, often absent, filiform. Leaves numerous, alternate on the stolons, up to 20 mm. long; segments capillary, glabrous, simple or usually dichotomously forked from or near the base, each segment sometimes again 1–3 or rarely more times forked. Ovary globose; style short but distinct; stigma lower lip semi-orbicular, upper short, ± obsolete. Capsule globose, 2–4 mm. in diameter, dehiscing into 2 lateral valves. Filaments linear; anther-thecae ± confluent. Seeds rather few (± 20–40), imbricate on the smooth placenta, lenticular with a broad irregular wing, 1–1.6 mm. in total diameter, outer surface smooth to verrucose; hilum prominent; testa-cells small, irregular, indistinct. Small to medium sized, annual or perennial, affixed or suspended aquatic. Rhizoids absent or present, filiform, a few cm long, 0.2-0.4 mm thick, bearing a few very short, dichotomously divided branches with curved, papillose, apically obtuse segments; sometimes the rhizoids with an indefinite apical transformation as a stolon. Stolons usually numerous, much branched and often mat-forming, filiform, terete, glabrous, up to 20 cm long or longer, 0.2-1 mm thick, sometimes somewhat thicker and spongy. Leaves numerous, 0.5-1.5 cm long, ± uniform, the primary segments 2 (or sometimes reduced to 1), sparsely dichotomously branched into up to 8 (but usually not more than 4) ultimate segments, or sometimes 1 branch of each dichotomy replaced by a trap, the ultimate segments capillary, slightly flattened, laterally entire or sparsely denticulate, the apex of the segments and of the lateral teeth when present setulose. Traps rather few, lateral on the primary to penultimate segments, ovoid, stalked, 1-2.5 mm long, the mouth lateral with 2 dorsal, setiform, branched appendages and further dorsal, lateral and ventral setae, the internal glands 2- and 4-armed, the arms narrowly ovoid, with apex acute, 10-40 µ long, 1.5-6 times as long as wide, the quadrifids with both pairs ± equally slightly divergent. Inflorescence erect, emergent, (or sometimes submersed, much reduced and with cleistogamous flowers), 1-20(-30) cm long, solitary or frequently 2 or more arising together or in succession; peduncle filiform, terete, glabrous, ± straight, 0.4-0.8 mm thick. Scales usually 1, sometimes 2, at or near the middle of the peduncle, similar to the bract, or frequently absent. Bracts basifixed, semi-amplexicaul, approximately semicircular, with apex rounded, truncate or obscurely dentate, c. 1 mm long, minutely glandular, obscurely nerved. Bracteoles absent. Flowers (1-)2-6(-12), the raceme axis short to elongate; pedicels erect or spreading, filiform, terete, 0.2-3 cm long. Calyx lobes subequal, broadly ovate or circular with apex rounded, 1-3 mm long. Corolla yellow, often with reddish-brown nerves, 4-25 mm long; upper lip broadly ovate to almost circular, usually ± obscurely to distinctly 3-lobed; lower lip limb usually slightly smaller, circular to transversely elliptic, the base with a prominent, rounded, bilobed swelling, the apex rounded, entire; palate densely pubescent with simple unicellular hairs; spur narrowly conical or narrowly cylindrical from a conical base with apex obtuse or ± obscurely bifid, a little shorter than to distinctly longer than the lower lip, the distal part externally, sparsely, minutely stipitate-glandular. Filaments curved, 1-1.5 mm long, the anther thecae ± confluent. Ovary globose; style distinct; stigma lower lip transversely elliptic, the upper lip obsolete. Capsule globose, 2-3 mm in diameter, the wall membranous, tardily laterally bivalvate. Seeds imbricate on the placenta, thickly lenticular, approximately circular in dorsal view, 0.8-1 mm in diameter, with a relatively broad, thick wing, the wing produced to form a rounded lobe on that part of the margin which, in situ on the placenta, is nearest the apex of the capsule, the rest of the margin shallowly and irregularly dentate, the ventral surface smooth or sometimes with multicellular, obtusely conical processes, the testa cells small, slightly elongate, with the anticlinal boundaries sunken, the periclinal walls smooth, convex, but with the central area collapsing and becoming concave. Flowering more or less throughout the year when conditions are suitable. Submerged, aquatic annual or perennial to 20 cm. Leaves sparsely dissected. Flowers in an ebracteolate, emergent raceme, yellow, lips subequal, broadly ovate, palate obscuring mouth, spur ± as long as or longer than lower lip. Fruits erect. Aquatic or subaquatic herb. Leaves with a few segments, 5-15 mm long. Flowers: bracteoles 0, inflorescence 1-4-flowered, without whorl of inflated leaf-like organs; corolla with spur well developed, 4-25 mm long, not hairy outside; yellow, often with brown or reddish nerves; Oct.-Jun. Traps ovoid, 1.0-1.5 mm long. Submerged, aquatic annual or perennial to 20 cm. Leaves sparsely dissected. Flowers in an ebracteolate, emergent raceme, yellow, lips subequal, broadly ovate, palate obscuring mouth, spur ± as long as or longer than lower lip. Fruits erect. Aquatic or subaquatic herb. Leaves with a few segments, 5-15 mm long. Flowers: bracteoles 0, inflorescence 1-4-flowered, without whorl of inflated leaf-like organs; corolla with spur well developed, 4-25 mm long, not hairy outside; yellow, often with brown or reddish nerves; Oct.-Jun. Traps ovoid, 1.0-1.5 mm long. Shallow still or slowly flowing water, or in mud in ditches, pools, lakes, river backwaters, bogs, swamps and marshes. Also frequently in deep water but not flowering unless supported on floating mats of living or dead vegetation. Usually at low altitudes but ascending to 2500 m alt. Boggy or shallow water. Dry Grassland. Shallow water and mud by ditches, streams and lakes. Boggy or shallow water. Dry Grassland. Shallow water and mud by ditches, streams and lakes. Pan tropical, extending northwards into eastern and western Canada and U.S.A., Spain and Portugal, North Africa, Israel, China and Japan and southwards to Argentina, S. Africa, Australia and New Zealand, also as an occasional introduction into other European countries and a common weed of aquaria and botanic gardens throughout the world. America: Canada (Nova Scotia, New Brunswick, Quebec, Ontario, [Manitoba], British Colombia), U.S.A.! (absent only in the great plains and Rocky Mountains states i.e. N & S Dakota, Nebraska, Iowa, Montana, Wyoming, Colorado, New Mexico, Arizona, Utah, Idaho, Nevada, and also Alaska, but in Hawaii), Mexico (Coahuila, Durango, Tamaulipas, San Luis Potosi, Jalisco, Tabasco, Michoacan, Mexico, Puebla, Chiapas, Campeche, Vera Cruz, Yucatan, and very probably elsewhere), Guatemala, El Salvador, Honduras, Belize, Nicaragua, Costa Rica, Panama, Bahamas, Cuba, Jamaica, Haiti, Dominican Rep., Puerto Rico, Antigua, Guadaloupe, Marie Galant, Dominca, Martinique, Trinidad!, Venezuela (Sucre, Monagas, Delta Amacuro, Anzoategui, Guarico, Miranda, Bolivar, Amazonas, Apure, Barinas, Portugesa, Cojedes, Carabobo, Falcon, Aragua, Yaracuy, Merida, Zulia), Colombia (Santander, Cundinamarca, Cauca, Amazonas), Guyana, Surinam, French Guiana, Brazil (Para, Amazonas, Maranhão, Ceara, Piaui, Pernambuco, Bahia, Goias, Mato Grosso, Minas Gerais, Espirito Santo, Guanabara, Rio de Janeiro, Sao Paulo, Sta. Catarina, Parana, Rio Grande do Sul), Ecuador, Peru, Bolivia, Paraguay, Uraguay, Argentina (Chaco, Jujuy, Salta, Tucuman, Corrientes, Misiones, Santa Fe, Cordoba, Buenos Aires). Europe: England (introduced), Portugal, Spain, Hungary (introduced). N. Africa: Morocco, Algeria, Egypt. Tropical and S. Africa: Senegal, Mali, Guinea-Bissau, Guinée, Sierra Leone, Liberia, Côte d’Ivoire, Burkina, Ghana, Nigeria, Niger, Cameroun, Central African Republic, Chad, Gabon, Congo, Zaïre, Sudan, Ethiopia, Uganda, Kenya, Rwanda, Tanzania!, Zambia, Zimbabwe, Moçambique, Angola, Namibia, Botswana, Transvaal, Natal, Cape Province, Madagascar, Mauritius. Asia: Israel, India (Rajasthan, Uttar Pradesh, Maharashtra, Madhya Pradesh, Andhra Pradesh, Karnataka, Tamil Nadu, Kerala, W. Bengal, Meghalaya, [Nicobar Islands]), Nepal, Ceylon, Burma, Thailand, Vietnam, Peninsular Malaysia, Sumatra, Java, Borneo, Philippines, Sulawesi, New Guinea, China (Yunnan, Guanxi, Guangdong, Hong Kong, Hunan, Henan, Fujian, Taiwan), Japan (southern), Palau. Australasia: Australia (Western Australia!, New South Wales, Queensland!, Northern Territory!), New Zealand (possibly introduced), New Caledonia. Uitenhage and throughout the tropics. Uitenhage and throughout the tropics. Inflorescence erect, 2–35 cm. high, solitary or often several arising in succession from the fascicle of stolons; flowers (1)2(6); peduncle straight, smooth and glabrous; scales absent or more usually 1 at or above the middle of the peduncle, similar to the bracts; bracts basifixed, semi-circular, semi-amplexicaul, more or less 1 mm. long; bracteoles absent; pedicels filiform, erect, (2)6–12(30) mm. long. Filaments linear; anther-thecae more or less confluent. Ovary globose; style short but distinct; stigma inferior lip semi-circular, superior short, more or less obsolete. Calyx lobes subequal, circular to broadly ovate, 1–3 mm. long. Corolla yellow, often with brown or reddish nerves, 4–25 mm. long; superior lip circular, 2–3 times as long as the upper calyx lobe, usually shallowly and obscurely 3-lobed; inferior lip usually shorter and narrower than the superior, circular, usually entire, rarely emarginate; palate much raised and bigibbous; spur conical to narrowly cylindrical, shorter than to almost twice as long as the inferior lip, the apex usually bearing a few shortly stalked glands. Seeds rather few (more or less 20–40), imbricate on the smooth placenta, lenticular with a broad irregular wing, 1–1.6 mm. in total diam., distal surface smooth to verrucose; hilum prominent; testa-cells small, irregular, indistinct. Capsule globose, 2–4 mm. in diam., dehiscing into 2 lateral valves. Aquatic or subaquatic herb. Rhizoids few from the base of the peduncle, often absent, filiform. Stolons usually up to more or less 10, fasciculate at the base of the peduncle, filiform, terete, up to 20 cm.×1 mm., sometimes slightly inflated. Traps usually numerous, replacing one of the leaf segments at a fork, ovoid, 1–1.5 mm. long, stalked; mouth lateral, with 2 dorsal, long, usually copiously branched hairs and a few ventral shorter simple hairs. Leaves numerous, alternate on the stolons, up to 20 mm. long; segments capillary, glabrous or sparsely setulose, simple or usually dichotomously forked from or near the base, each segment sometimes again 1–3 or rarely more times forked. Filaments linear; anther-thecae more or less confluent. Ovary globose; style short but distinct; stigma inferior lip semi-circular, superior short, more or less obsolete. Calyx lobes subequal, circular to broadly ovate, 1–3 mm. long. Corolla yellow, often with brown or reddish nerves, 4–25 mm. long; superior lip circular, 2–3 times as long as the upper calyx lobe, usually shallowly and obscurely 3-lobed; inferior lip usually shorter and narrower than the superior, circular, usually entire, rarely emarginate; palate much raised and bigibbous; spur conical to narrowly cylindrical, shorter than to almost twice as long as the inferior lip, the apex usually bearing a few shortly stalked glands. Seeds rather few (more or less 20–40), imbricate on the smooth placenta, lenticular with a broad irregular wing, 1–1.6 mm. in total diam., distal surface smooth to verrucose; hilum prominent; testa-cells small, irregular, indistinct. Capsule globose, 2–4 mm. in diam., dehiscing into 2 lateral valves. Aquatic or subaquatic herb. Rhizoids few from the base of the peduncle, often absent, filiform. Stolons usually up to more or less 10, fasciculate at the base of the peduncle, filiform, terete, up to 20 cm.×1 mm., sometimes slightly inflated. Traps usually numerous, replacing one of the leaf segments at a fork, ovoid, 1–1.5 mm. long, stalked; mouth lateral, with 2 dorsal, long, usually copiously branched hairs and a few ventral shorter simple hairs. Leaves numerous, alternate on the stolons, up to 20 mm. long; segments capillary, glabrous or sparsely setulose, simple or usually dichotomously forked from or near the base, each segment sometimes again 1–3 or rarely more times forked.General Information
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Morphology
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Habitat
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Distribution
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Manual de Plantas de Costa Rica
General InformationFlora of China @ efloras.org
General InformationBrazilian Flora 2020 project - Projeto Flora do Brasil 2020
MorphologyFlora de Nicaragua
General InformationManual of Vascular Plants of Northeastern US and Canada
General InformationPlants Of the World Online Portal - FTEA
Morphologye-Flora of South Africa
MorphologyFlora Zambesiaca - descriptions
Morphology