Herbs erect, 1-2 m tall, densely pubescent and glandular. Petiole 3-5 cm; leaf blade broadly ovate, 10-18 × 4-15 cm, membranous or felty, minutely tomentose, base rounded or obtuse, asymmetric, margin subentire, sinuate, or irregularly dentate, apex acute; veins 7-10 pairs. Flowers erect. Pedicel 1-5 cm. Calyx cylindric, 8-10 × 2-3 cm; lobes narrowly deltate, 1-2 cm, sometimes unequal. Corolla greenish proximally, white at apex, 15-18 cm; limb 7-10 cm in diam.; lobes mucronate at apex. Filaments ca. 5.5 cm; anthers 1-1.7 cm. Capsules deflexed, globose or ovoid, 3-4 cm in diam., densely armed with slender, subequal prickles, pubescent with white hairs, irregularly dehiscent at apex, subtended by remnants of persistent calyx. Seeds numerous, brown, discoid-reniform, 3-5 mm in diam. Fl. and fr. Jun-Sep. Herb to subshrub 0.5-1 m tall, with strong fetid odor, branching appearing pseudodichotomous; branches herbaceous, nearly cylindrical, soft pilose, sometimes purple-tinged. Leaf blades 6-16 x 3-11.5 cm, irregular but usually with ovate outline, chartaceous, soft-pubescent, the apex shortly acuminate, obtuse, acute, or acuminate, the base very unequal, rounded-acute, the margins deeply sinuate or lobed (the lobes rounded); petioles 3-7 cm long, pubescent. Flowers erect, sweet-scented. Calyx green, nearly tubular but widened toward the base, 5-9.5 cm long, the lobes ca. 1 cm long, ovate-triangular; corolla white funnel-shaped, the limb pentagonous, slightly reflexed, with acute to acuminate angles, the accessory lobes acuminate, the tube 10-13 cm long, narrowed toward base; stamens included, the anthers white or cream-colored, connivent; style white, the stigma yellow, bilobed, included. Capsule on reflexed peduncle, with ovoid outline, to 3 cm long, densely covered with sharp spines to 1 cm long, opening along 4 longitudinal valves, from top to bottom, the reflexed calyx base irregularly lobed. Seeds numerous, reddish brown, ca. 4 mm long. Plant up to 130 cm tall, branched, dense pubescent-villous. Leaves 7-18 x 4.5-12 cm, broadly ovate, repand to sinuate-dentate, base oblique to cuneate. Petiole shorter than lamina. Pedicel 10-25 mm long, reflexed in fruit, tomentose with brownish hairs. Calyx 7.5-10.5 cm long, 1/2-2/3rd the corolla length, 5-lobed, slightly inflated toward the base; basal portion persistent and strongly reflexed in fruit; lobes unequal, triangular or ovate-lanceolate, acuminate, up to 22 mm long. Corolla tube 15-17 cm long; limb 6-8.5 cm broad, 10 (-12) lobed; lobes broad triangular, acute or acuminate. Anthers 9-11 mm long, white. Capsule globose, 4-5 cm broad, dense spiny, nodding; spines acicular, 5-10 mm long. Seeds 4.5-5 mm long, reniform, compressed, brown, minutely reticulate, rugose-lineate on the back. Folha: indumento pubescente(s); margem(ns) inteira ou sinuosa(s). Flor: cálice(s) maior(es) que 4 cm compr.; corola cor branca. Fruto: pedúnculo(s) reflexo(s); pericarpo espinescente(s). Semente: cor bege. Hoja: indumento pubescente(s); margen(es) entera o sinuosa(s). Flor: cáliz mayor(es) que 4 cm largo; corola color blanca. Fruto: pedúnculo(s) reflexo(s); pericarpo espinescente(s). Semilla: color beige. Leaf: indumentum pubescent; margin entire or sinuose. Flower: calyx bigger than 4 cm long; corolla colour white. Fruit: peduncle reflexed; pericarp spinescent. Seed: colour beige. Annual or perennial herb, up to 1 m or more high. Pedicels pubescent, up to 20 mm long. Calyx 95-100 mm long. Corolla up to 200 mm long. Fruits covered with numerous narrow, stiff spines up to 15 mm long. Fruits and spines pubescent. Flowers pink or lavender. Common in recently disturbed areas such as roadsides and waste grounds. Emmaus (A1991), Great Cmz Bay (A2362). Also on St. Croix, St. Thomas, and Tortola; southeastern United States to South America, including the West Indies, introduced throughout the tropics. Herbs erect, 1-2 m tall, densely pubescent and glandular. Petiole 3-5 cm; leaf blade broadly ovate, 10-18 × 4-15 cm, membranous or felty, minutely tomentose, base rounded or obtuse, asymmetric, margin subentire, sinuate, or irregularly dentate, apex acute; veins 7-10 pairs. Flowers erect. Pedicel 1-5 cm. Calyx cylindric, 8-10 × 2-3 cm; lobes narrowly deltate, 1-2 cm, sometimes unequal. Corolla greenish proximally, white at apex, 15-18 cm; limb 7-10 cm in diam.; lobes mucronate at apex. Filaments ca. 5.5 cm; anthers 1-1.7 cm. Capsules deflexed, globose or ovoid, 3-4 cm in diam., densely armed with slender, subequal prickles, pubescent with white hairs, irregularly dehiscent at apex, subtended by remnants of persistent calyx. Seeds numerous, brown, discoid-reniform, 3-5 mm in diam. Fl. and fr. Jun-Sep. Plant up to 130 cm tall, branched, dense pubescent-villous. Leaves 7-18 x 4.5-12 cm, broadly ovate, repand to sinuate-dentate, base oblique to cuneate. Petiole shorter than lamina. Pedicel 10-25 mm long, reflexed in fruit, tomentose with brownish hairs. Calyx 7.5-10.5 cm long, 1/2-2/3rd the corolla length, 5-lobed, slightly inflated toward the base; basal portion persistent and strongly reflexed in fruit; lobes unequal, triangular or ovate-lanceolate, acuminate, up to 22 mm long. Corolla tube 15-17 cm long; limb 6-8.5 cm broad, 10 (-12) lobed; lobes broad triangular, acute or acuminate. Anthers 9-11 mm long, white. Capsule globose, 4-5 cm broad, dense spiny, nodding; spines acicular, 5-10 mm long. Seeds 4.5-5 mm long, reniform, compressed, brown, minutely reticulate, rugose-lineate on the back. Folha: indumento pubescente(s); margem(ns) inteira ou sinuosa(s). Flor: cálice(s) maior(es) que 4 cm compr.; corola cor branca. Fruto: pedúnculo(s) reflexo(s); pericarpo espinescente(s). Semente: cor bege. Hoja: indumento pubescente(s); margen(es) entera o sinuosa(s). Flor: cáliz mayor(es) que 4 cm largo; corola color blanca. Fruto: pedúnculo(s) reflexo(s); pericarpo espinescente(s). Semilla: color beige. Leaf: indumentum pubescent; margin entire or sinuose. Flower: calyx bigger than 4 cm long; corolla colour white. Fruit: peduncle reflexed; pericarp spinescent. Seed: colour beige. Hoja: indumento pubescente(s); margen(es) entera o sinuosa(s). Flor: cáliz mayor(es) que 4 cm largo; corola color blanca. Fruto: pedúnculo(s) reflexo(s); pericarpo espinescente(s). Semilla: color beige. Leaf: indumentum pubescent; margin entire or sinuose. Flower: calyx bigger than 4 cm long; corolla colour white. Fruit: peduncle reflexed; pericarp spinescent. Seed: colour beige. Terrícola Erva Subarbusto Subarbusto Annual or perennial herb, up to 1 m or more high. Pedicels pubescent, up to 20 mm long. Calyx 95-100 mm long. Corolla up to 200 mm long. Fruits covered with numerous narrow, stiff spines up to 15 mm long. Fruits and spines pubescent. Flowers pink or lavender. Weed of waste places. Native to North America. Common in recently disturbed areas such as roadsides and waste grounds. Emmaus (A1991), Great Cmz Bay (A2362). Also on St. Croix, St. Thomas, and Tortola; southeastern United States to South America, including the West Indies, introduced throughout the tropics. Herb to subshrub 0.5-1 m tall, with strong fetid odor, branching appearing pseudodichotomous; branches herbaceous, nearly cylindrical, soft pilose, sometimes purple-tinged. Leaf blades 6-16 x 3-11.5 cm, irregular but usually with ovate outline, chartaceous, soft-pubescent, the apex shortly acuminate, obtuse, acute, or acuminate, the base very unequal, rounded-acute, the margins deeply sinuate or lobed (the lobes rounded); petioles 3-7 cm long, pubescent. Flowers erect, sweet-scented. Calyx green, nearly tubular but widened toward the base, 5-9.5 cm long, the lobes ca. 1 cm long, ovate-triangular; corolla white funnel-shaped, the limb pentagonous, slightly reflexed, with acute to acuminate angles, the accessory lobes acuminate, the tube 10-13 cm long, narrowed toward base; stamens included, the anthers white or cream-colored, connivent; style white, the stigma yellow, bilobed, included. Capsule on reflexed peduncle, with ovoid outline, to 3 cm long, densely covered with sharp spines to 1 cm long, opening along 4 longitudinal valves, from top to bottom, the reflexed calyx base irregularly lobed. Seeds numerous, reddish brown, ca. 4 mm long.General Information
Source: [
Source: [
Source: [
Morphology
Source: [
Source: [
Distribution
Source: [
Flora of China @ efloras.org
General InformationFlora of Pakistan
General InformationBrazilian Flora 2020 project - Projeto Flora do Brasil 2020
Morphologye-Flora of South Africa
MorphologyMemoirs of the New York Botanical Garden
Distribution