The World Flora Online

Plants 30-60 cm tall. Rhizome erect or obliquely ascending, short, stout, scaly; scales pale brown, ovate or ovate-lanceolate, membranous, dentate. Fronds caespitose; stipe stramineous, as long as lamina, longitudinally grooved, with ovate, dentate scales; lamina deltoid or ovate-oblong, 20-30 cm, 15-16 cm wide at lower part, bipinnate-pinnatifid; pinnae 12 or 13 pairs, oblique, basal pair largest, shortly stalked, upper ones gradually shortened, connected by wing at base, lowest ones deltoid, 9-10 × 4-5 cm, base subtruncate, shortly stalked, apex acuminate; pinnules 8 or 9 pairs, basal pair largest, upper ones gradually shortened, connected by broad wing, basal pair opposite or subopposite, deltoid, 2.5-3 cm, base 1-1.5 cm wide, shortly stalked; acroscopic pinnules smaller than basiscopic ones, nearly at right angle to costae, basiscopic one parallel to pinna rachis or slightly oblique, pinnatifid; segments 4 or 5 pairs, basal pair asymmetrical, basiscopic one smaller than others, oblique, acroscopic one oblong, 8-10 × ca. 2 mm, almost parallel to costa, aristate. Veins impressed adaxially, raised abaxially, pinnate, simple, extending into each tooth. Sori terminal on apex of lateral veins, in 2 rows along main vein of pinnules; indusia pale brown, membranous, with an incised margin, often deciduous at maturity.

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Leaves monomorphic, dying in winter, 15--75 × 10--30 cm. Petiole 1/4--1/3 length of leaf, scaly at least at base; scales scattered, tan. Blade light green, ovate-lanceolate, 2--3-pinnate-pinnatifid, herbaceous, not glandular. Pinnae ± in plane of blade, lance-oblong; basal pinnae lanceolate-deltate, slightly reduced, basal pinnules usually longer than adjacent pinnules, basal basiscopic pinnule longer than basal acroscopic pinnule; pinnule margins serrate, teeth spiny. Sori midway between midvein and margin of segments. Indusia lacking glands. 2 n = 164.

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Rhizome horizontal, short-creeping; lvs deciduous, essentially glabrous except for the chaffy, concolorous pale brown scales that beset at least the lower part of the petiole and sometimes also part of the rachis; petiole mostly a quarter to a third as long as the blade; blade 2–5+ dm, a third to half as wide, broadest a little below the middle, bipinnate- pinnatifid to tripinnate, with mostly 10–15 pairs of pinnae below the pinnatifid tip; pinnae (at least the lower ones) mostly oblique, the basal pinnule of the lower side of the lowest pinna longer than the one next to it and to twice (thrice) as long as the subopposite upper pinnule; ultimate segments finely spinulose- toothed; sori midway between the midvein and the margins; 2n=164. Moist or wet woods and swamps; interruptedly circumboreal, in Amer. s. to S.C., Ark., and Wash. (D. spinulosa; D. austriaca var. s.)

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Çokyıllık. 30-120 cm boyunda bitkiler. Rizom kuvvetli, kısa, 10 × 3-4 cm, yatık-yükselici veya yatay, çok pullu, ölü yaprakların kalıntıları ile örtülü. Fırontlar 30-120 cm, 2 teleksi; petiyol az çok aya ile eşit uzunlukta, nispeten ince; pullar yumurtamsı ilâ yumurtamsı-mızraksı, yüreksi, ortası koyu olmayan saç görünüşlü, aşağıda koyu kahverengi, daha sık, yukarıda daha soluk; aya 15-60 × 10-25 cm, dış hatları dikdörtgensi-mızraksı ilâ mızraksı veya nadiren yumurtamsı- dikdörtgensi, parlak yeşil ilâ sarımsı-yeşil, nadiren alt yüzeyde az sayıdaki salgılı tüyler hariç çıplak. Telekler 15-20 çift, kısa saplı, alttaki çiftler karşılıklı veya yarı-karşılıklı, üsttekilerin çoğu almaşlı, aşağıdaki çift genellikle bir üsttekinden çok az kısa, uçtaki telek mızraksı gibi; telekçik dikdörtgensi-yumurtamsı, dekürent, tabandaki derince teleksi, diğer telekçiklerin her biri uca doğru az derince kesik, en sondaki parçalar yumurtamsı-dikdörtgensi teleksi dişli, dişler dikencikli-mukronat, eğik; eksen seyrek, bizsi pullu. Soruslar çok sayıda, çoğunlukla telekçiğin ikincil damarlarının çatallarında çifter çifter; indizyum yuvarlak-böbreksi, kenarları parçalanmamış, dalgalı veya az dişli, nadiren birkaç saplı salgılı.

Blätter 15–90 cm lang, hellgrün, kahl, Stiel so lang wie die Spreite oder länger, spärlich mit einfarbig blassbraunen Spreuschuppen besetzt. Spreite 2,5–4mal so lang wie breit, nach dem Grund kaum verschmälert, doppelt gefiedert. Fied

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7–8. Saure Waldböden, Waldsümpfe, Hochmoore, kollin-subalpin. CH.

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